THE SNAKES OF EUROPE BY G A BOULENGER LL.D., D.SC, PH.D F.R.S., F.Z.S WITH FOURTEEN PLATES AND FORTY-TWO FIGURES IN THE TEXT METHUEN & GO LTD 36 ESSEX STREET W.G " ' LONDOl First Published in igi3 PREFACE is no work in the English language with the Reptiles of Europe I have therefore endeavoured to supply this desideratum, so THERE dealing Snakes are concerned, by drawing up in a concise form an account of what is known of their far as the characters, their distribution, and their life-histories Professor Sordelli, of Milan, having kindly acceded to my request to reproduce some of the beautiful figures drawn by him for the work published in collaboration with the late Professor Jan under the title of" Iconographie Generale des Ophidiens," I have been able to supplement descriptions with illustrations which leave nothing to be desired from the A few drawings have point of view of accuracy my been made specially for this book by Mr J Green I have further to acknowledge the permission given by the Trustees of the British Museum, the India Office, figures and the Zoological Society, to reproduce a few from previous publications of which I am the author In order to render this little book more useful, the account of the Snakes of Europe has been preceded THE SNAKES OF EUROPE vi by an Introduction summarizing what Snakes generally I this known is of have purposely avoided overburdening a work of kind, which aims at concision, with bibliographi- cal references and synonymic lists I am sure my readers will be thankful for being spared this display of erudition Whenever I have had to compile, and to trespass on ground that is not my own, I have been careful to draw only from the writings of the most trustworthy authorities The descriptions of the species are based on the collection in the British Museum, which has been considerably increased since the publication of the Catalogue (1893-1896) I of Snakes have also had access to Monsieur F Lataste's rich private collection, now under my care, and Dr R Gestro has kindly entrusted to me for study the collection of Italian Snakes in the Genoa Museum I am indebted to Dr L W Sambon for the chapter on Parasites, which he has written at my request To all who have helped me I beg to tender my hearty thanks G A B CONTENTS INTRODUCTION CHAPTER I II I'AGE Definition and Classification External Characters — Integument - - i - - III Coloration - - IV Skeleton Dentition Poison Apparatus - - - - - - - -53 V VI - Poisons VII VIII IX - - Nervous - Viscera - — Different - - Organs of Reproduction tion Development - ; 62 - - - -77 XI Parasites - - 82 - - - - - - - - - - - 107 - - - - - 118 Man - - - XII Distribution XIII Snakes in 73 Pairing; Oviposi- - ; X Habits 40 of - Organs - 29 Kinds - - System — Sense - - Relation to 91 133 SYSTEMATIC ACCOUNT OF THE SNAKES OF EUROPE First Family: TYPHLOPIDjE Genus TYPHLOPS, Schneider - Typhlops vermicularis, Merrem Blind-Snake - - - — The Greek 144 144 vii viii THE SNAKES OF EUROPE Second Family : BOID^E PAGE Genus ERYX, Daudin Eryx jaculus, Linnaeus— The - - - 147 - 147 Javelin Sand-Boa Third Family: COLUBRID^E Genus TROPIDONOTUS, Kuhl Tropidonotus natrix, Linnaeus— The - - - Snake, or Ring-Snake Tropidonotus - - 152 Lauren ti — The 160 - Tropidonotus viperinus, Water-Snake perine Genus ZAMENIS, Wagler - tessellatus, Tessellated Water-Snake Latreille — The - Vi- - - - — — — - - vandi's Snake - - Coluber leopardinus, ard Snake 13 - Coluber dione, Pallas — The Dione Snake — Laurenti The ^EsculaColuber longissimus, pian Snake 12 Coluber - — Bonaparte The scalaris, Schinz — The Ladder Snake - - - Snake - - Coronella girondica, Smooth Snake - - - Daudin — The - 181 182 185 187 Leop- Genus CORONELLA, Laurenti — The Smooth 14 Coronella austriaca, Laurenti 15 170 - Genus COLUBER, Linn^us quatuorlineatus, Lacepede— Aldro9 Coluber 11 165 Zamenis gemonensis, Laurenti The European 170 Whip-Snake Zamenis dahlii, Fitzinger Dahl's Whip-Snake 177 Zamenis hippocrepis, Linnaeus The Horseshoe' \Whip-Snake 179 - 10 152 Grass- - 191 194 196 197 Southern 202 CONTENTS ix PAGE Genus CONTIA, Baird and Girard - 205 - 205 - - 207 Genus MACROPROTODON, Guichenot — The 18 MACROPROTODON CUCULLATUS, I Geoffroy 212 16 Ccelopeltis - False Smooth Snake - Genus TARBOPHIS, Fleischmann 19 - Hermann monspessulana, Montpellier Snake - Contia modesta, Martin— The Dwarf Snake Genus CGELOPELTIS, Wagler 17 - — The 208 - - - 213 - - - 216 Tarbophis fallax, Fleischmann — The Cat-Snake Tarbophis iberus, Eichwald — The Caucasian Cat- 217 - 219 - 221 Vipera ursinii, Bonaparte— Orsini's Viper Vipera renardi, Christoph Renard's Viper Vipera berus, Linnaeus The Northern Viper, or Adder Vipera aspis, Linnaeus— The Asp Viper Vipera latastii, Bosca Lataste's Viper Vipera ammodytes, Linnaeus The Sand-Viper, 221 Snake - Fourth Family: VIPERID^E Genus VIPERA, Laurenti 21 22 23 - - - - — -— - - 24 25 26 — or 27 - — — Long-Nosed Viper Vipera lebetina, Linnaeus Viper, or Kufi - - - - - - - 230 239 247 249 The Blunt-Nosed - - Genus ANCISTRODON, Palisot de Beauvois 28 Ancistrodon halys, Pallas— Pallas's Pit-Viper Index - 227 - - - 257 - 261 - 262 - 265 LIST OF PLATES FACING PAGE PLATE I II Typhlops vermicularis, Eryx jaculus 144 Tropidonotus natrix and vars, cettii and PERSA -152 - III IV - Tropidonotus tessellatus, VAR AUROLINEATUS - HIPPOCREPIS - IX X XI l6o - Z 170 dahlii, - - 176 Coluber quatuorlineatus and var sauro- Coluber longissimus, C VAR QUADRILINEATUS VIII - - MATES, C DIONE VII - Zamenis gemonensis, var caspius, Z VI viperinus and Zamenis gemonensis and vars persica and VIRIDIFLAVUS V T - - - 82 88 leopardinus and - Coluber scalaris- - - CORONELLA AUSTRIACA - - - - - 194 - 196 CORONELLA GIRONDICA, CONTIA MODESTA COZLOPELTIS MONSPESSULANA, M ACROPROTODON - 202 CUCULLATUS, TARBOPHIS IBERUS, T FALLAX 208 - VlPERA URSINII, V RENARDI, V BERUS XIII VlPERA ASPIS, V LATASTII XII - - 220 - 240 XIV VlPERA LEBETINA, HALYS V AMMODYTES, ANCISTRODON XI 250 General Literature Leaders of Religion II C BEECHING With Edited by Crown Cardinal Newman Svo 2s Portraits net each volume Thomas Chalmers R H Hutton Mrs Oliphant Secon-j Edition John Wesley J H Overton Bishop Wilberfokce G Cardinal Manning John Knox John Howe A H Charles Simeon W W Lancelot Andrewes Daniell Hutton Augustine of Canterbury R F Horton Thomas Ken W H William Laud Cutt.s Third Edition A J Mason R M and A Latimer W Bishop Butler Walter Lock Four.', i Augustus Jessop Thomas Cranmer T Hodgkin Hutton Edition John Donne F A Clarke George Fox, the Quaker John Keble E L C G Moule MacCunn Second Edition F R L Ottley Second Edition J Carlyle A Spooner The Library of Devotion With Introductions and (where necessary) Notes Smalt Pott Svo, The Confessions Eighth Edition of St 2s.; 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