Begun NUMBER in 1895 JULY 358 Neogene Paleontology in the Northern Dominican Republic 20 Holoplanktonic Mollusks (Gastropoda: Heteropoda and Thecosomata) by Arie W Janssen Palcontological Research Institution 1259 Trumansburg Road Ithaca, New York, 14850 U.S.A 23, 1999 PALEONTOLOGICAL RESEARCH INSTITUTION Officers Shirley K Egan President John C Steinmetz Thomas E Whiteley Henry W Theisen Howard P Hartnett First Vice-President Second Vice-President Secretary Treasurer Warren Director D Allmon Trustees Mary M Shuford Constance M Soja John C Steinmetz Carlton E Brett William L Crepet J Thomas Dutro, Jr Shirley K Egan Howard P Hartnett Harry G Lee Amy Gregory P Wahlman Thomas E Whiteley McCune R Megan Peter B Stifel Henry W Theisen D Shay Trustees Emeritus Harry A Leffingwell Robert M Linsley Samuel T Pees Edward B Picou, Jr James E Sorauf Raymond Van Houtte William P S Ventress BULLETINS OF AMERICAN PALEONTOLOGY and PALAEONTOGRAPHICA AMERICANA Warren D Allmon Editor Reviewer for this issue Paula M Mikkelsen A list of titles in both series, and available numbers and volumes may be had on request Volumes 1-23 oi Bulletins of American Paleontology are available from Periodicals Service Company, 11 Main St., Germantown, New York 12526 USA Volume of Palaeontographica Americana has been reprinted by Johnson Reprint Corporation, 1 Fifth Ave., New York, NY 10003 USA Subscriptions to Bulletins of American Paleontology are available for US $150 per year (individual or institution) plus postage Issues are available and priced individually Numbers of Palaeontographica Americana are priced individually for additional information, write or call: Paleontological Research Institution 1259 Trumansburg Road Ithaca, NY 14850 USA (607) 273-6623 FAX (607) 273-6620 www.englib.comell.edu/pri @ This paper meets the requirements of ANSI/NISO Z39.48-1992 (Permanence of Paper) f\/7(^-7 ^u(Qtms of Begun NUMBER in 1895 ^t^^ JULY 358 Neogene Paleontology in the Northern Dominican Republic 20 Holoplanktonic Mollusks (Gastropoda: Heteropoda and Thecosomata) by Arie W Janssen Paleontological Research Institution 1259 Trumansburg Road New York, 14850 U.S.A Ithaca, 23, 1999 ISSN 0007-5779 ISBN 0-87710-448-4 Lihraiy i>f Confitess Catalog This publication is Card Number: 99-64085 supported in part by a Corporate Membership from Exxon Exploration Company Note: Beginning with issue number 356, Bulletins of American Paleontology is no longer designating volumes The journal will continue to publish approximately 2-4 issues per year, each of which will continue to be individually numbered Printed in the United Slates of America Allen Press, Inc Lawrence, KS 66044 U.S.A CONTENTS Page Abstract Resumen Introduction Acknowledgements Abbreviations Used in the Text Biostratigraphy and Correlations Early Miocene Late Miocene Assemblages Pliocene Assemblages Conclusions Systematic Palaeontology Order Heteropoda 11 Family Atlantidae Order Thecosomata 11 13 Family Limacinidae 13 Family Cavoliniidae 15 Subfamily Creseinae 15 Subfamily Cuvierininae 19 Subfamily Clioinae Subfamily Cavoliniinae 21 22 References Cited 28 Plates 33 Index of Holoplanktonic Mollusk Taxa 39 LIST OF ILLUSTRATIONS Page Text-figure Chrono-, Range Neotype of litho- and biostratigraphical mterpretations (after Saunders et ul mollusk species Gurabo section (Quoy and Gaimard, 1827) LIST 10 19 OF TABLES Table and overall distribution of holoplanktonic chart of holoplanktonic mollusk species in the Ri'o Stylicilu suhiila 1986), Measurements (in mm) of Ciivieriiui aslesanti (Rang 1827)? Measurements of Ccivnlinia gypsorum (Bellardi 1873) Dimensions of complete specimens of Cavolinia mexicana (Collins, 1934) Page 20 23 25 NEOCENE PALEONTOLOGY IN THE NORTHERN DOMINICAN REPUBLIC 20 HOLOPLANKTONIC MOLLUSKS (GASTROPODA: HETEROPODA AND THECOSOMATA) Arie W Janssen National Museum 2300 RA of Natural History (Palaeontology Department, Cainozoic Mollusca), P.O Leiden, The Netherlands, and 12, Triq il-Hamrija, Xewkia VCT 110, Box 9517, Gozo, Malta ABSTRACT Holoplanktonic gastropod faunas from the northern Dominican Repubhc Neogene contained History Museum at rolitndahi (Gabb, 1873), and 17 euthecosomatous pteropods; inflata (d'Orbigny in the collections of the Natural Basel (Switzerland) comprise two species of Heteropoda; Allanta conlifdniti.s Gabb, 1873 and Prolallanta Limacina (Striolimacina) imilans (Gabb 1873), L (Striolimacina) 1836) Limacina sp indet Creseis aciciila (Rang, 1828), Hyalocylis striata (Rang, 1828), Styliola suhula (Quoy and Gaimard 1827) Cuviehna astesana (Rang, 1829)?, Cuvierina sp., Clio cuspidata (Bosc, 1802)?, C pyramidata Linne, 1767 forma kmceokiia (Lesueur, 1813), Clio sp., Cavolinia gypsorum (Bellardi 1873) C aff gypsorum, C me.xicana (Collins, 1934) C cf irideiilalci (Niebuhr 1775), Diacria trispinosa (de Blainville, 1821 1873) recorded in the literature from the same area but not represented in the ) and Edilhinella sp Edithinella iiiuhilala present collection, is (Gabb, included on the basis of the The number of holoplanktonic gastropod species from the Dominican Republic is thus 20 The name Slrioliinacina is introduced to replace Phmorhella Gabb 1873a non Haldemann 1843 (Mollusca) From the Baitoa Formation (Ri'o Yaque del Norte section), of assumed late Early Miocene age, a single pteropod (Edilhinella sp.) was recorded The Late Miocene Cercado Formation (Ri'o Gurabo: Globorotalia hiimerosa Zone, Ri'o Cana; no biozone indicated) and some samples lacking lithostratigraphical data, of ? Late Miocene age (Ri'o Mao) yielded nine species, among which only Cavolinia gypsorum is indicative of a Late Miocene (Tortonian-Messinian) age Other species are known only locally type material or have longer ranges Specimens from the late Early Pliocene Gurabo Formation {Globorotalia margaritae Zone; Rio Gurabo) and the Mao Adentro Limestone (G inurgarilae Zone; Ri'o Canal, the Early to middle Pliocene Mao Formation (G nuirgarilaelmioceiiica Zone; Ri'o Gurabo section), and a sample lacking lithostratigraphical data, of ? late Early Pliocene age (Rio Yaque del Norte section) yielded 10 holoplanktonic mollusk species So far as they could be identified to species these invariably belong to taxa ranging through the entire Pliocene, and still at least occur today RESUMEN La asociacion de Gasteropodos holoplanctonicos colecciones del Museo de del Neogeno del Norte de la Repiiblica Dominicana que se encuentra en las Historia Natural de Basilea (Suiza), esta compuesta por dos especies de Heteropoda: Atlanta cordiformis Gabb 1873 y Protallanta rotuiulata (Gabb 1873) y 17 pteropodos eutecosomatos: Limacina imitans (Gabb 1873) L inflata (d'Orbigny 1836) Limacina sp indet Creseis acicula (Rang 1828) Hyalocylis striata (Rang 1828) Styliola subiila (Quoy and Gaimard, 1827), Cuvierina astesana (Rang, 1829)?, Cuvierina sp Clio cuspidata (Bosc 1802)'? C pyramidata Linne 1767 forma lanceolata (Lesueur 1813), Clio sp., Cavolinia gypsorum (Bellardi, 1873), C aff gypsorum C me.xicana (Collins, 1934), cf tridentata (Niebuhr 1775), Diacria trispinosa (de Blainville, 1821) y Edithinella sp Se incluye para su estudio la especie Edithinella undulata (Gabb 1873) descrita de la misma region pero ausente en la coleccion de dicho museo basandose en el material tipo Asi se conoce un total de 20 especies de Gasteropodos holoplanctonicos del Neogeno de la Repiiblica Dominicana Se introduce el nuevo nombre Striolimacina para el genero Planorhella Gabb 1873a non Haldemann 1843 (Mollusca) En la Formacion Baito (el corte del Rio Yaque del Norte), considerada de tener una edad Mioceno Final tardio se encontrado un solo pteropodo: Edithinella sp La Formacion Cercado de edad Mioceno Final (Ri'o Gurabo: Globorotalia huinerosa Zona; Ri'o Cana: no biozona indicada) y algunas muestras sin precision litoestratigrafica consideradas de tener la misma edad (Ri'o C Mao) han librado nueve especies Entre ellas Cavolinia gypsorum es la unica especie caracteristica del Mioceno Final (Torton- iense-Messiniense) Las otras especies o bien son endemicas o tienen una distribucion estratigrafica larga Formacion Gurabo (Rfo Gurabo) y de la Caliza de Mao Adentro (Rio Cana) de edad Plioceno Antiguo Formacion Mao de edad Plioceno Antiguo-Mediano (G margaritae/miocenica Zona, Rio Gurabo) y una muestra considerada de tener una edad Plioceno Antiguo tardio (Ri'o Yaque del Norte) han producido 10 Moluscos holoplanctonicos que en cuanto sido posible su determinacion a nivel especi'fico pertenecen sin excepcion a taxones un rango temporal desde el Plioceno Antiguo hasta la actualidad Especimenes de la tardio (Globorotalia margaritae Zona), de la INTRODUCTION Dominican Republic, between 1978 and 1980 Geographic and stratigraphic information on the various The material of holoplanktonic moUusks studied in paper originally comprised almost 150 samples, collected from outcrops in the Cibao Valley, in the this was supplied by Saunders et al (1986), to which reference is made, With a few exceptions, samples here referred to are sections Bulletin 358 Museum One Abbreviations Used taxon recorded in the literature from the Dominican Republic, but not represented in the material before The following abbreviations housed me is in the Naturhistorisches included The at Basel on which it was some species repand was received original material based, as well as type material for resented in the collection survives, on loan from American ANSP vation Other samples, however, comprise excellently GMHU Geological MHNP Musee which apparently were sorted view of techniques were special collecting apno shells, holoplanktonics, the composition of the material material does not include new was obtained on national d'Histoire naturelle, Paris, Naturhistorisches NNM ment, Basel, Switzerland; National Museum of Natural History (Inverden, RGM The Netherlands; Museum National ca), Leiden, The Netherlands (formerly: Rijksmuseum van Geologic en Mineralogie); USNM National Museum ogy, Washington all I wish to thank Dr Peter Jung for entrusting the material to thermore I thank all (NMB) me for description Furwho made information tory, J Collier (National Smithsonian Institution, Museum of Natural His- Department of Paleobi- DC, U.S.A.), Professor George M Gary Rosenberg (Academy of Natural ology, Washington Davis and Dr Sciences, Department of Malacology, Philadelphia, U.S.A.), Dr P Lozouet (Musee national d'Histoire naturelle, Laboratoire de Biologic des Invertebres Marins et Malacologie, Paris, France); Dr Saul Rothmann (Geological Museum, The Hebrew University, JeiTi- salem, Israel), and Professor H Shibata (Department of Geology, College of General Education, Nagoya University, Nagoya, Japan) I am grateful to Dr Peter Jung (NMB), Dr Irene Zorn (Geologische Bundesanstalt, Vienna, Austria) and to my colleague Dr C F Winkler Prins (NNM) for critical reading of the manuscript, the last named and Dr M Freudenthal (NNM) supplied the "Resumen" Dr Maria del Carmen Perrilliat Institute de Geologia, Mexico DF, Mexico) kindly furnished information on stratigraphy and literature John W M Jagt (Venlo, The Netherlands) improved the English, ( as usual SEM gelen (NNM) made the Goud (NNM) Jeroen final version prepared the and Mrs Mieke van Enprinted them and reduced and repro- micrographs of Plate (NNM) height of shell; W width of D dorso-ventral diameter of shell shell; Biostratigraphy and Correlations The holoplanktonic molluscan material from the Cibao Valley, as studied in the present paper, was found to comprise 19 species, of which belong to the Heteropoda, and 17 to the Euthecosomata Of these II (57.9%) could be identified at species level, two are identified with a query, and six remain in open nomenclature Material was available from 31 stations, distributed 16 stations), Rio Cana Rio Mao (five stations), and Ri'o Yaque del Norte (two stations) The distribution of these samples and their species contents over the various chrono-, litho- and biostratigraphical units, as specified in Saunders et al (1986) are shown in Text- over four sections: Rio Gurabo (eight figure ( stations), I The holoplanktonics from the Rio Gurabo section offer the most interesting details and the most complete picture by far Biostratigraphically these samples are assigned to three planktonic Foraminifera zones, viz the Late Miocene Glohorotalia humerosa Zone, the late Early Pliocene Glohorotalia margaritae Zone, and the Early to middle Pliocene Globorotalia mar- garitae/miocenica Zone Dirk van der Marel of Text-figure DC, U.S.A.; H colleagues or collection material in their care available to me: Mr Frederick of Natural History, Smith- sonian Institution, Department of Paleobiol- Acknowledgments of of Natural History (Pa- laeontology Department, Cainozoic Mollus- rare or ill-known species First Museum, Geology Depart- tebrate Department, Recent Mollusca), Lei- still taxa, but in various cases valuable additional information University, Je- NMB quite interesting and deserves to be studied in detail The Museum, Hebrew France; carefully collected sieving residues In plied, nor special attention paid to the occurrence of is Philadel- rusalem, Israel; ding planes, usually in a rather poor state of preser- the fact that are used: phia, U.S.A.; institutions Quite a number of samples consist of pieces or small from the Te.xt The Academy of Natural Sciences, slabs of sediment with specimens preserved on bed- preserved isolated in duced the drawings 1, From the Ri'o Cana planktonic mollusks section samples yielded holo- One of these is assigned a late Early Pliocene Globorotalia margaritae Zone age, whereas the remaining seven lack biozone details, age being indicated as "? Late Miocene" their Dominican Neocene Holoplanktonic Mollusks: Janssen The same is true for the five Rio Mao samples in which holoplanktonics were found They are not zoned, and their age hkewise is '"? Late Miocene'" Two samples are available from the Rio Yaque del Norte section, both yielding but a single species The upper one is not zoned; its age is assumed to be late quite Formation of Santa Rosa, Vera Cruz, Mexico (as Limacina elevata Collins) The same or a very closely related species with the same type of microsculpture has also been recorded from the Early Pliocene (Zanclean) of the Mediterranean area (Italy and France) These occurrences will be evaluated in a paper in prep- Early Pliocene The lower sample, from the late Early Miocene Baitoa Formation, finally, yielded a single aration specimen and lacks biostratigraphical data as from the same well Cavolinia mexicana, interestingly, was introduced locality Santa Rosa Mexico This in species has also been recorded from the Late Mio- Early Miocene Assemblages The sole specimen of Edithinella sp known ently belongs to an undescribed species and fore of The little appar- Of much there- is use for both biozonation and correlation related Edithinelki imdulota also based cene-middle Pliocene of Japan (zones bata and Ujihara, 1990) on a sin- section This is the that species It was described from of northern resented in the material studied for this paper Edithi- p was based on a single "middle Miocene" specimen from the Panama Canal Zone, in addition to this occurrence the species is known only from three Miocene European localities nella cahbbetnta (Collins, 1934) type Italy, first (Shi- occurrence of Cavolinia gypsoriim found in two samples of the Rio known specimen, has previously been recorded from the "Miocene" of the Dominican Republic, without any further information This species is not repgle interest also is the N 18-20) Gurabo extra-Mediterranean record of "Messinian" the but a nannoplankton analysis of the showed its age to be Tortonian (Janssen, 1995, Other records of this species from Europe are considered to be Messinian This is a good first- lot 102) still order correlation across the Atlantic, and, in only certain indication fact, the among holoplanktonic Mollus- of no use for correlations; the species, is indeed a Late Miocene assemblage samples from the Gurabo section yielded specimens here referred to as Cavolinia aff gypsorum in both cases co-occurring with gypsorum Very similar, if not identical, material has been recorded from the Late Miocene-Middle Pliocene of Japan (N18-20) (Shibata and Ujihara, 1990) A direct comparison with the Japanese specimens will be necessary to settle the identity of both In two samples of the Gurabo section specimens of a Cuvierina species were found, here indicated as C astesana'l They differ morphologically slightly from sam- the admittedly highly variable Pliocene individuals of (Janssen, 1995) It too is not represented in the present material ca that this Two C Late Miocene Assemblages In the Rio Gurabo section the Late Miocene assem- blage yielded seven species, one of which {Limacino infiata) has been recorded from the Middle Miocene of Australia and Turkey (Janssen, 1990), as well as from many younger assemblages, and Its occurrence range makes is it is still therefore not surprising and incidentally, has not been found extant its in the Pliocene long C astesana ples Of the other six species (one heteropod and five euthecosomatous pteropods) two are exclusively known from Caribbean area (Protatlanta rotimdata and Limacina imitans) L imitans occurs in one sample only, P rotundata is present in four samples (of the seven) P rotimdata, a heteropod originally described from "Santo Domingo", has never been recorded from elsewhere, but it might be closely related or even conspecific with the Plio/Pleistocene Protatlanta kakegawaensis, from Japan Furthermore, various related species are present in the RGM collections from localities of Miocene and Pliocene age in the Mediterranean area (unpublished) Limacuui imitans is an interesting species from a taxonomic point of view It was likewise described from "Santo Domingo" and is also known from the Early Pliocene (Zone N19; Akers, 1972) Agueguex- from the type area northern in they are apparently older than typical C Italy As astesana they might represent a precursor species, which cannot any of the various Miocene Cuvierina species, however, from the Mediterranean area The samples from the Rio Cana section indicated as "'? Late Miocene" yielded six species, five of which are also known from the Gurabo Late Miocene, viz Protatlanta rotundata, Limacina imitans, Cavolinia aff gypsorum, and mexicana Four gypsorum, samples from this interval, however, also contain Diacria trispinosa, a species that in the Rio Gurabo section occurs exclusively in the Pliocene portion D trispino.ui, however, is also known from Late Miocene sediments in Italy (Janssen, 1995) and thus its occurclearly be related to C C rence in the Cana section its absence in the is more easily explained than Late Miocene of the Gurabo section In the five samples species were found from the Rio One of these is Mao section four the heteropod At- Bulletin 358 Llthostratigraphy Chronostratjgrapby E it a - Species Biostratlgraphy I a No s i f a :§ i s •3 a = i2 « 3 I- F :§ :§ I I H Rio Gurabo section 15833 15832 15829 • • 15823 15828 15827 15993 15851 15854 15906 15907 15904 15900 15903 15913 15914 Rio Cana section 17023 16835 16836 16837 16838 16844 16856 16875 Rio Mao section 16915 16923 16927 16922 16932 Rio Yaque del Norte section 17293 17288 — Text-figure I Chrono-, litho- and biostratigraphic interpretations of and overall distribution of holoplanktonic mollusk species lanta cordiformis, originally described from Santo mingo and not recorded from elsewhere since Do- Two Limacina imitans and Cavolinia gypsorum are also known from the Late Miocene interval of the Rio Gurabo and Rio Cana sections The fourth species species, again Diacria trispinosa, present in the Cana section, but absent along the Rio Gurabo Miocene tran- tlie Neogene ot the northern The few Europe, From the distribution of holoplanktonic mollusks it Miocene samples from the Rio and the "? Late Miocene" ones from clear that the Late Gurabo section, the Rio Cana and Mao sections (as specified in Text- figure 1) are correlative Saunders er 1986) possibilities of long-distance correlation to especially the occurrence of Cavolinia gypsorum, could indicate a Tortonian/Messinian age of these intervals Pliocene Assemblages Only from samples of Zone Gurabo section is more or less from the Pliocene available: four the Rio substantial material sect (after i.e is is Dominican Repubhc late Early Pliocene Globorotalia margari- from the Early to mid- dle Pliocene Globorotalia margaritae/miocenica Zone tae age, and five samples yielded a total of 10 species, all euthecosomatous pteropods These biozones have five pteropod species Bulletin 358 32 Mizunami Fossil Museum, voL I, pp 73-91, tigs., pis 1880 23-26 e fossili nel terreno terziario italiano aggiunte e cor- Shibata, H., and T Ishigaki 1981 Mu- English summary) Bulletin of the Mizunami Fossil vol pp 55-70 hgs tabs., vol 6, pp Torelli, L., 1981 12-13 pis Shizukawa Groups of Shimobe, Yamanashi Prefecture, no 13 pp 39-55 Mizunami Museum, Fossil tab., figs., pis vol Ja- 1990 tab I 13, 1990 fig., pis zuoka Prefecture, Japan Venus, and time, and of the Mizunami Fossil 1974 Museum, bioseries their Monographs 1921 Pteropodi del Miocene di S Nicola Varano (Gargano) 417-430, 20 Romana, vol in figs RC.A.L Rang & FL.A Souleyet, 1852 Histoire nade mollusques pteropodes Monographic compre- nant la West description de toutes les especes Bailliere, Paris, 1970 parts of Amsterdam), 375 pp 17 tabs., 366 pp., 246 Holkema Utrecht, 484 ries, pt (CD-ROM) pis the older Tertiary of Australia Trans- Australia, vol 9, pp 194-196, pi Zorn, 20 Leiden Museum, the Lcyden museum Notes from vol 29, pp 181-203, 306D, pp 37 figs., maps, 40 pis, of Semata The Jour- Tokyo, vol 19, no 220, pi (2) I 1991 The Pteropoda of no pp 1-8, figs., pi 1(1); vol 19, no 223, pp 17-24, actions and Proceedings of the Royal Society of South Tesch, J.J of 1-564 48-66 nal of the Geological Society of The pteropods of i-vii Pt Marginellidae to Helminthoglossi- Survey Professional Paper Yamakawa, G., and M Ishikawa 1912 Some pteropods from the Neogene Tate, R A systematic account of Tertiary Pteropoda (Gastropoda Euthecosomata) from Austria Contributions the and Quaternary Geology, pi vol 28, no 4, pp to Tertiary 95-139 20 tabs., 12 figs., 16 pis Tiberi, N 1878 Eulimidae the Republic of Haiti, pp 1-631, Pelagic Mollusca of the World World Biodiversity Se- in press results Publications Woodring, W.P., J.S Brown and W.S Burbank 1924 Geology of the Republic of Haiti Geological Survey of figs Spoel, S van der (ed.) Do- Panama Description of Tertiary mollusks (gas- 299-452, Pseudothecosomata, Gymnosomata and Heteropoda (Gas- the 89-154 Miocene mollusks from Bowden Jamaica dae) Geological figs tropoda) Bohn Scheltema and pp 40 pis Geology and paleontology of Canal Zone and adjoining tropods: (thesis University of 1908 Indies Noorduijn, Gorinchem J I, Gastropods and discussion of Euthecosomata, a group with remarkable developmental stages (Gastropoda, Pteropoda) geological reconnaissance of the Carnegie Institute, Washington 385, pp pp 1-86, 15 pis Spoel, S van der 1887 A Wiegniann, A.F.A., and J.F Ruthe 1832 Handbuch der Zoologie Luderig, Berlin, pp i-vi, 1-621 Woodring, W.P 1928 Contributions to the Geology and Palaeontology of the turelle 1976 et al (eds) Dominican Republic Geological Survey of minican Republic, Memoir 7, pp 1967 Jonio Conchiglie vol 10, nos 5-6 pp 93-1 12, Tertiary and Quaternary stratigraphic paleontology In: the figs Souleyet, F.L.A 1852 Mar T.W Vaughan no 7, pp 61-82, tabs., descrizione di due nuove specie Geologica 306— Vaughan, T.W., and W.P Woodring Sirna, G 1968 vol 49 no 4, pp pis pis Itoigawa and H — figs., Sui molluschi di alcuni saggi di fondo prelevati alle soglie del In: J figs Vatova, A 43-47 Shibata (eds) Faunas and floras of Cenozoic Japan distribution in space Sedimentary basins of Me- (ed.) Pteropods from the Sagara Group (Mio-Pliocene), Shi- 329 tabs., Cenozoic pteropod fauna of Japan FC Wezel raphy, Tecnoprint, pp 173-186, tabs 7-8 Middle and Late Pleistocene heteropods and pteropods from Chiba, Noto Peninsula and Kikaijima, Japan Bulletin of the Mizunami Fossil Museum, vol 10 pp 151169 /;;: Ujihara, A., H Shibata and T Saito Shibata, H., and A Ujihara 1983 Late Quaternary stratigraphy of the Sardinia Basin sedi- diterranean margins C.N.R Italian Project of Oceanog- Pteropods from the Late Neogene Nishiyatsushiro and pan Bulletin of the 5-49 and G Buccheri ments Shibata, H., T Ishigaki and A Ujihara 1986 Bollettino della Societa Malacologica Italiana rezioni Heteropodous and pteropodous biostratigraphy of Cenozoic strata of Chubu Province, Japan (in Japanese with seum, Cefalopodi, pteropodi, eteropodi viventi nel Medilerraneo Mollusques marins d"Italie Cephalopodes, pteropodes, du terrain tertiaire d'ltalie Annales de la Societe Malacologique de Belgique, vol 13, no 2, pp 52-84 heteropodes vivant de la Mediterranee et fossiles 1997 Planktonic gastropods from the Early Messinian of the Heraklion Basin (Crete, Greece) Submitted to Contributions to Tertiary pp 31-45, and Quaternary Geology, figs., 4pls vol 34, no 1- PLATES Bulletin 358 34 Explanation of Plate Page Figure Limacinu imitans (Gabb, 1873) la-f NMB H 17616, Sta 15900 13 Gurabo; Cercado Formation (Late Miocene, Glohnrntalia humerosa Zone); la frontal view, microsculpture barely visible (bar length = 100 p,m); lb frontal view of body whorl, showing peripheral spiral and backwards diverging microsculpture below and above periphery (bar length = 100 jJLm); Ic peripheral view of body whorl showing dense Ri'o diverging microsculpture (bar length = 100 \Lm): sculpture fading out towards the suture (bar length fading of microsculpture (bar length sculpture and new = 10 |xm); body whorl near apertural margin, showing diverging view of body whorl near aperture, showing irregular close up of early body whorl, showing growth lines intersected by microId apical side of = 100 If jim); le oblique sculptural elements inserting in apertural direction (bar length = 10 jjim) Bulletins of American Paleontology, Bulletin 358 Plate Bulletins of American Paleontology, Bulletin 358 9d Limacina imitans Plate Dominican Neocene Holoplanktonic Mollusks: Janssen 35 Explanation of Plate Page Figure 1-2 Atlanta conlifonms Gabb 1873 11 la-d Lectotype (Pilsbry, 1922), Id left lateral 2a-d NMB H ANSP 2896; Santo Domingo (Miocene); apical view; lb frontal view; Ic umbilical view; la view; X 25 17617, Sta 16923; Rio 2a apical view; 2b frontal Mao at mouth of Arroyo Bajon; level E no formation name (age unclear ? Late Miocene); view; 2c umbilical view; 2d rear view; X 25 3-4 Protatlanta rotundata (Gabb 1873) 3a-e Holotype, view; 4a-c X NMB H ANSP 2891; Santo at mouth of Arroyo Bajon; level E; 4a apical view; 4b frontal view; 4c umbilical view; X X 50 Cercado Formation (Late Miocene Glohorotalia 12.5 Protatlanta smileyetl (Smith, 1888) 5a-c NNM collection umbilical view; 6-9 3a apical view; 3b umbilical view; 3c frontal view; 3d right lateral 25; 3e oblique frontal view (with protoconch in upright position); 17618 Sta 15903; Rio Gurabo, humerosa Zone); 11 Domingo (Miocene); Limacma X Canary 12 Islands, Sta 2.114, Tydemann Expedition 1977 12.5 imitans (Gabb 1873) 6a—d Neolectotype 13 (hie design.) 6d right lateral view; x ANSP 2895; Santo Domingo (Miocene); Formation (Miocene); NMB H 6a apical view; 6b frontal view; 6c umbilical view; 25 7a-d Limacina inflata Collins, 1934 (non d'Orbigny 1836); 8a-d (Recent); 5a apical view; 5b frontal view; 5c USNM 483145; Rio Mao Sta 17619, Sta 16927; Rio Mao 1979 at mouth of Arroyo Bajon level E; frontal view; 8b apical view; 8c oblique frontal view; 8d umbilical view; 9a-e Holotype of Limacina etevata Collins, 1934; USNM X no formation name (Late Miocene); Limacina lOa-b inflata NMB H 645189; Santa Rosa, Vera Cruz, Mexico (Miocene); 9a apical view; Limacina lla-d X 25 (d'Orbigny 1836) 14 17620, Sta 15903; Rio Gurabo 1978; Cercado Formation (Late Miocene, Glohorotalia humerosa Zone); lOa apical view; 10b frontal view showing opaque subperipheral 8a 25 9b right lateral view; 9c oblique frontal view; 9d frontal view; 9e umbilical view; 10 8525 (Vaughan and Cooke); Cercado x 25 7a frontal view; 7b right lateral view; 7c apical view; 7d umbilical view; inflata (d'Orbigny USNM belt, the thickening is visible at the apertural margin; x 25 1836)? 645188; Santa Rosa, Vera Cruz, Mexico (Miocene); same specimen illustrated in Collins, 1934, pi 7, Limacina inflata): la apical view; lib frontal view; lie oblique frontal view; lid umbilical view; X 25 I 14 figs 3-5 (as Bulletin 358 36 Explanation of Plate Figure 1-2 Page Cre.wix acicuhi (Rang 1828) 1-2 NMB H 15 Mao 17621 Sta 15829: Rio Gurabo; Formation (Early to middle Pliocene Glohomtalia margaritae/iniocenica Zone): x 50 3-5 Hyalocylis striata (Rang 1828) NMB H 16 Mao Formation (Early to middle Pliocene Glohorotalia margaritae/iniocenica damaged specimen of Clio pyramidala f lanceolalci is visible in the lower left) 17622, Sta 15829; Ri'o Gurabo: Zone): x (a 4a-b Holotype of Hyalocylis haitensis (Collins 1934): Sta 9574 (Miocene); USNM 371905; Port-au-Prince, near Petionville 4a ventral view: 4b left lateral view: 5a-d Illustrated paratype of Hyalocylis haitensis (Collins, 1934): x USNM 371905: same x section; 5b ventral view: 5c lower transverse section: 5d right lateral view; Haiti, US Geol Surv 7.5 locality as 4a-b; 5a upper transverse 7.5 Hyalocylis striata (Rang, 1828)? 16 Lectotype (Janssen, 1995) and illustrated specimen of Tentaciilites creiaceiis Blanckenhorn, 1889; GMHU 2100: Nisib, Turkey: yellowish-white limestone (Miocene ? Vindobonian): also illustrated in Blanckenhorn 1899 pi 22, fig (as Avnimelech 1945, fig (as Clio cretaceiiin (Blanckenhorn): x (same slab also contains "Vaginella rotiinihilu" Blanckenhorn (no syntype) (= Vaginella sp.?) "Balantiiiin flahellifonne" Blanckenhorn (syntype) (= Propeanutssiiiin sp.) and "Cavolinia" sp (= indet foraminifer?) and Tentaciilites cretaceiis) 7-9 Styliola in (Quoy and Gaimard, 1827) siihiila 18 7a-b Holotype of Styliola sulcifera Gabb, 1873; ANSP 2893; Santo Domingo ("•Miocene"), also illustrated in Pilsbry, 1922, and Collins, 1934, pi 9, figs 9, 9a-10 (as Styliola sulcifera): 7a upper transverse of specimen in Pilsbry, 1922): X 12.5 8a-b NMB H 17623, Sta 15854: Ri'o Gurabo: Gurabo Formation (Early Pliocene Glohorotalia nuirgaritae Zone): 8a ventral view, X 12.5: 8b apical shell part (tip damaged), x 50 text-fig (2 figs., as Styliola sulcifera) section: 7b ventral 9a-b NMB H view (dotted line is outline 17524 Sta 15851: Ri'o Gurabo: Gurabo Formation (Early Pliocene Glohorotalia nuirgaritae Zone): 9a adapical view: 9b ventral view; x 12.5 10 Cuvierina aslesana (Rang, 1827)7 lOa-c NMB H 17625 Sta 15907: 19 Ri'o Gurabo: Cercado Formation (Late Miocene, Glohorotalia hunierosa Zone): view: 10b ventral view: 10c right 1 Cuvierina sp 20 NMB H 17626, Sta 15828: Ri'o Gurabo: ventral view (?), 12 view: x lateral 10a adapical X 12 (surrounding matrix Mao Formation (Early to middle Pliocene, Glohorotalia nuirgaritae Zone); omitted) Clio cuspidala (Bosc 1802)? 12 NMB H 21 17627 Sta 15829 Zone): ventral view, x NMB H 14a-f NMB H Gurabo: f Formation (Early to middle Pliocene, Glohorotalia niargaritae/miocenica to middle Pliocene, Glohorotalia margaritae/iniocenica lanceolala (Lesueur, 1813) 17628, Sta 15829: Ri'o Gurabo: Zone); dorsal view, x Mao 13-17 Clio pyramidala (Linne 1767) 13 Ri'o Mao 21 Formation (Early 17629, Sta 15993 Ri'o Gurabo; Gurabo Formation (Early Pliocene Glohorotalia margaritae Zone): 14a dorsal view: 14b left lateral view; 14c ventral view, 14d adapical view, x 12.5; I4e protoconch, frontal view; 14f idem, left view (tip missing), x 25 15a-c Holotype of Cleodora howdenensis Collins, 1934 USNM 645194; Bowden Bowden Beds, lower zone (Late Pliocene, but indicated "Miocene"): 15a dorsal view: 15b transverse section at aperture; 15c ventral view; x 16 Specimen illustrated by Collins (1934 pi 12 fig 7: "may also belong to C howdenensis") USNM 371908: Jacmel Haiti: US(GS) Sta 9604 (Pliocene): dorsal view X 10 lateral 17 Specimen sp cf (Miocene?): dorsal 18 Vaughan and Woodring 1922 p 166) as Carolina sp and by Collins 1934 p 202) as Cleodora 371907; El Mores Monte Cristi Dominican Republic: USGS Sta 8783: Yaque Group view, x referred to by howdenensis; ( ( USNM Clio sp 21 Protoconch NMB H 17630 Sta 15829 Ri'o Gurabo; miocenica Zone); 18a transverse section Mao Formation (Early at aperture; 18b frontal to view: 18c middle Pliocene, Glohorotalia nuirgaritae/ lateral view; x 25 Bulletins of American Paleontology, Bulletin 358 Plate ^ 15a ij' -•: Clio pyramidata f lanceolata *^' ; U V- !^W: m Bulletins of American Paleontology, Bulletin 358 Cavolinia gypsorum Plate ^~* 3a %:4S ;? Dominican Neocene Holoplanktonic Mollusks: Janssen 37 Explanation of Plate Page 22 Figure 1-3 Cavoliniu gypsoruin (Bellardi, 1873) la-c NMB H view; 17631, Sta 15903 Ri'o Gurabo; Cercado Formation (Late Miocene Globorolalia humerosa Zone); lb right lateral Interlocking view; mechanism of Ic ventral view; x dorsal shell pan, la dorsal left NMB H side; 17632 Sta 16837 Ri'o Cana; Cercado Formation (probably Late Miocene); x 12.5 3a-c, Isolated ventral shell part, illustrated by Collins (1934 Mao, Cavolinia aff 4a-c 8525; Cercado Formation ("Middle Miocene"); Sta gypsoruin (Bellardi 1873) an NMB H sp pi figs 8-9 as Cavolinia sp indet.); 3a left lateral USNM 371904; view; 3b ventral view; 3c adapical view; x Ri'o 23 nov.? 17633 Sta 15907 Ri'o Gurabo; Cercado Formation (Late Miocene Globorolalia humerosa Zone); 4a dorsal view; 4b right lateral view; 4c ventral view; X 24 5-8 Cavolinia me.xicana (Collins 1934) 5a-d Holotype, USNM 645206; Santa Rosa, Vera Cruz, Mexico; US(GS) Sta 9995 ("Middle Miocene"); 5b left lateral view; 5c adapical view; 5d ventral view; x 5a dorsal view; and NMB H 17635 (fig 7), Sta 16837 Ri'o Cana; Cercado Formation (probably Late Miocene); 6a-7a dorsal views; 6b-7b right lateral views; 6c-7c ventral views; 7d adapical view; x 8a-b Aberrant specimen with oblique furrows on ventral side; NMB H 17636 Sta 15913 Ri'o Gurabo; Cercado Formation (Late Miocene Globorolalia humerosa Zone); 8a ventral view; 8b right lateral view; X 6a-c, 7a-d NMB H 17634 (fig 6) Bulletin 358 38 Explanation of Plate Page Figure 1-3 CavoUuia cf iridentata (Niebuhr, 1775) NMB H Isolated ventral shell pan; Fragment of dorsal X NMB H Formation (Early to NMB H side; NMB H 17638 locality data as in fig x NMB H 17639, Sta 15828 Rio Gurabo; Mao Formation 1; X 26 x (surrounding matrix f Mao Formation (Early bisiilcaia to bisulcata Gabb, 1873; NMB H Miocene); 7a dorsal view; 7b ventral view; x Specimen of f bisulcala Gabb; middle Pliocene GloboioutUa imuguritae/iniocenica Zone): Gabb 1873; ANSP 2894; and Collins (1934 pi 6b adapical view; 6c right lateral view; 6d ventral view; f view; omitted) illustrated in Pilsbry (1922, text-fig 4, figs.) 7a-b Specimen of left lateral 6a-d Holotype of Diacria trispinosa NMB H x S Domingo, Dominican Republic ("Miocene"); also 18-20 (copied from Pilsbry); 6a dorsal view; figs 17642, Sta 16915 Rio Mao; at mouth of Arroyo Bajon, level E (? Late Mao Formation (Early x (surrounding matrix not drawn) 17643, Sta 15823 Rio Gurabo; roralia margaritae/miocenica Zone); dorsal view, (Early to Middle Pliocene Glohorolaliu 17641, Sta 15823 Rio Gurabo; dorsal view, middle Pliocene Globonitalia inargaritae/mioceiuca Xone): 17640 Sta 16844 Rio Cana; Cercado Formation (probably Late Miocene); 4a dorsal view; 4b 4c ventral view; Mao margaritae Zone); dorsal view, x 4-8 Diacria trispinosa (de Blainville 1821) 4a-c 25 17637 Sta 15823 Rio Gurabo; to middle Pliocene, Globo- 27 Edithinella sp 9a-c NMB H 17644, Sta 17288 Rio Yaque del Norte; Lopez section; Baitoa Formation (? late Early Miocene); 9a adapical view; 9b left lateral view; 9c dorsal view; X Bulletins of American Paleontology, Bulletin 358 Plate INDEX OF HOLOPLANKTONIC MOLLUSK TAXA acicula acicula Creseis 16 Clio (Creseis) acicula 15 acicula forma acicula Creseis 16 Clio (Styliola) Lamherti 18 acicula, Cleodora 15 Clio acicula 15 acicula, Cleodora (Creseis) 15 Clio bowdenensis 2! acicula, Clio 15 Clio braidensis 21 acicula, Clio (Creseis) 15 Clio cretaceum acicula, Creseis 16; PI 3, figs 9, 15, 1-2 acicula, Creseis acicula forma acicula, Creseis acicula acicula, Vaginella 17 21 22 Clio cuspidata 16 Clio cu.spidatal 16 clio helicina 15 Clio pyramidata 9, 21; PI 3, fig 12 13 21, 22 13-17 15 Clio pyramidata forma lanceolata anguslicoslata, Cavolinia 26 Clio sp Anomia Clio striata columnella, Cuvieria 19 20 aciculala Hyalaea 7, 19, 20; Pi fig 10 columnella Cuvierina 11 cordiformis, Atlanta astesana?, Cuvierina aslesana, Cuvierina 7, Atlanta Atlanta cordifnnnis 7, 1; PI 2, figs 1-2 Atlanta peroni 11 Creseis acicula acicula Atlanta rotunda 11 Creseis acicula forma acicula Atlanta souleyeti 16 Balantium flahelliforme Balantium undulatum bisulcala, Cavolinia [Diacria) bisulcata, Diacria trispinosa forma 27; PI 5, bowdenensis, Cleodora Creseis acicula 15, 16; PI 3, figs 16 1-2 16 16 Creseis spinifera 19 18, Creseis striata 16 11 cretacea, Praehyalocylis 17 cretaceum, Clio 17 18 cretaceus, Praehyalocylis 27 cretaceus Tentaculites 26 cuspidata'?, Clio fig bowdenensis Cleodora 1-2 PI 2, figs 1; 11 26, 27 bisulcata, Diacria 27 Allantidea Balantium 7, 15, 11,14 11,12 18 16 Creseis Atlanta inflata Atlanta rotundata 9, 21, 22; PI 3, figs 9, 21; PI 3, fig 26 20 Iridentata 17 16-18 12 21; PI fig 21, 22 cuspidata Clio Cuvieria columnella 21 Cuvierina 19 7, 19, 20 19; PI 3, fig 10 21 Cuvierina astesana bowdenensis Clio 21 Cuvierina columnella braidensis Clio 21 Cuvierina globosa Camptoceratops prisca 11 Cuvierina grandis 20 7, 28 Cuvierina inflata 21, 26 Cuvierina intermedia Cavolina iridentata 25 Cuvierina paronai Cavolinia 22 Cuvierina sp Cavolinia (Diacria) bisulcata 26 Cuvierina tubulata sp cf caribbeana Edithinella Cavolina sp Cavolinia aff gypsorum 7, 23; PI 4, fig Cavolinia cf iridentata 9, 25; PI 5, figs Cavolinia globulosa Cavolinia grandis 9; Cavolinia gypsorum 3, Cavolinia mexicana Cavolinia n 7-9, 22-25; 3, 7, 9, sp Cavolinia sp Cavolinia tridentata PI 3; figs 1-3 20 20 9, 20; PI fig 20 Diacria bisulcata 26, 26 27 Diacria digitata Diacria trispinosa 26 Diacria trispinosa forma bisulcata 1-3 Diacria trispinosa forma trispinosa digitata, 18 7-9, 26, 27; PI 5, figs 27; 4-7 PI 5, fig Edithinella 22 23 Edithinella caribbeana 26 Edithinella sp 15 Edithinella undulata 26 18 Diacria 22 23 9, 22, 21 Diacria 24 24, 25; PI 4; figs 5-8 Cleodora Cleodora [Balantium) 26 Cavolinia anguslicostata 3, 7, 27, 28 28 7, 7, 28; PI 5, fig 7, 27, 28 28 16 Edithinella varanica Cleodora (Balantium) undulatum 27 elevata, Cleodora (Creseis) acicula 15 elevala, Spiratella inflata Cleodora (Creseis) 16 Embolus Cleodora acicula 15 euphratensis Hyalocylis 16-18 Cleodora bowdenensis 21 flahelliforme Balantium 18 Cleodora sp Cleodora sp 21 globosa, Cuvierina 21 globulosa, Cavolinia striata 16 grandis, Cavolinia Cleodora suhula 18, 19 grandis, Cuvierina Cleodora Clio cf striata striata bowdenensis 17, 21 22, 28 Limacina 7, 13, 14 13 14 rostralis gypsorum, Cavolinia 24 9, 26 20 7-9, 22-25; PI figs 1-3 1 Bulletin 358 40 gypsorum, Cavolinia 23 24; PI 4, aft' haitensis, Hyalocylix fig hailensis Hyalocylis 16, Embolus rosi rails IS rotunda, Atlanta 17 rotundata, Atlanta hailensis, Praehyalocylis 17 rotundata, Protatlanta helicina forma rangii Limacina 11 rotundata Vaginella helicina, clio 13 souleyeti, Atlanta Hyalaea 15 souleyeti, Protatlanta aciciilata 14 II 7, 1, 12; PI 2, 11,12 figs 3-4 18 11 12; PI 2, fig, Hyalocylis 16, 18 sp Clio PI 3, fig 18 Hyalocylis euphratensis 16-18 ,sp., Cuvierina PI fig 18 sp Edilhinella 18 spinifera, Creseis Hyalocylis haitensis Hyalocylis ohtiisa 17, Hyalocylis striata 16-18; PI 3, figs Hyalocylis striatal 3-5 PI fig 16-17 Hyalocylix haitensis PI 5, fig 18, 19 Spiratella 13 Spiratella inflata elevata 13 Spirialis tertiaria 14 17 striata, Cleodora 16 6-9 striata, 16 13-15 striata Cleodora {Balantium) Cleodora (Creseis) imitans Valvatella 13 striata Clio 16 inflata elevata Spiratella 13 striata Creseis 1, 14 striata Hyalocylis PI 2, fig 10 striata, Hyalocylis striata striata, Hyalocylix Hyalocylix striata Limacina imitans 7-9, 13-15; PI 1, PI figs imitans, Planorhella inflata Atlanta inflata, Cuvierina inflata, Limacina inflata?, 7, 13-15; Limacina PI 2, fig 20 intermedia, Cuvierina kakegawaensis, Protatlanta Lamberti Clio 7, Styliola 18 Styliola Limacina Limacina elevata 14 Styliola suhula 7, mexicana, Cavolinia miiltispira 7, I, Styliola sidcifera 11 suhula, Cleodora 13-15; 6-9 PI 2, fig 10 PI 2, fig 15 3, 7, 9, 24, 25; PI 4, 14-15 figs 5-8 17 16, 17 19 3, 9, 18, 19 subula, Styliola tertiaria, 14, 15 14 tertiaria, Spirialis tridenlata, Anomia 26 tridentata Cavolinia 10 tridentata, Cavolinia cf 9, 22, 25 25; PI 5, figs trispinosa forma trispinosa, Diacria peroni, Atlanta II trispinosa Diacria Planorhella 14 trispinosa, Diacria trispinosa 13-15 7-9 16-18 Limacina trispinosa forma bisulcata, Diacria Planorhella imitans 19; PI figs 18 19 18 17, 18 sulcifera, Styliola 20 Hyalocylis 18, 19; PI figs.7-9 18 19 paronai Cuvierina obtiisa 17 Tentaculites cretaceus Plolophysops PI fig 16, 17 13, 14 PI 2, figs 3-5 18 Styliola striata PI PI 3, figs Rangiana 9,21,22 7-9, 13-15; 16-18; striata, Styliola lanceolata Clio pyramidata forma Limacina helicina forma rangii Limacina imitans Limacina inflata Limacina inflatal Limacina sp Limacina sp indet Limacina tertiaria 16 9, Hyalocylis striata?, 11, 12 (St^^liola) 16 27; PI 5, 26 1-3 fig 26 7-9, 26, 27; PI 5, figs forma tubulata Cuvierina 4-7 26 20 27, 28 Plotophysops multispira 10 uiutulata Edilhinella Praehyalocylis 18 undulata, Vaganella 27 Praehyalocylis crelacea 17 undulata, Vaginella 27 Praehyalocylis cretaceus 17 undulala, Vaginella aff 27 Praehyalocylis haitensis 17 iindulatum, Balantium 27 Camptoceratops 10 undulatum, Cleodora (Balantium) 27 Vaganella undulata 27 Vaginella 28 prisca, Protatlanta Protatlanta kakegawaensis Protatlanta rotundata Protatlanta souleyeti pyramidata forma lanceolata Clio pyramidata lanceolata, Clio pyramidata, Clio Rangiana, Styliola rcmgii Limacina helicina forma 7, 1, 1-13 11, 12 12; PI 2, figs 3-4 12; PI 2, fig 9, 21, 21, Vaginella acicula Vaginella aff undulata 7, 15 27 28 22 Vaginella rotundata 18 21 Vaginella sp 28 22 Vaginella undulata 27 18 Valvatella imitans 13 11 varanica, Edilhinella 28 ... numbers and volumes may be had on request Volumes 1-23 oi Bulletins of American Paleontology are available from Periodicals Service Company, 11 Main St., Germantown, New York 12526 USA Volume of Palaeontographica... Houtte William P S Ventress BULLETINS OF AMERICAN PALEONTOLOGY and PALAEONTOGRAPHICA AMERICANA Warren D Allmon Editor Reviewer for this issue Paula M Mikkelsen A list of titles in both series,... with issue number 356, Bulletins of American Paleontology is no longer designating volumes The journal will continue to publish approximately 2-4 issues per year, each of which will continue