Bulletins of American paleontology (Bull. Am. paleontol.) Vol 346

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>-3- OLUME ^u((ctins of "fAmcrxcan toioqs) Begun 107, in 1895 NUMBER 346 JUNE Brachiopods of the Onondaga Formation, Moorehouse Member (Devonian, in the Genesee Valley, Western Eifelian), New York by Howard R Feldman Paleontological Research Institution 1259 Trumansburg Road New York, 14850 U.S.A Ithaca, 15, 1994 PALEONTOLOGICAL RESEARCH INSTITUTION Officers President Vice-President J Thomas Dutro, John Jr C Steinmetz Henry W Theisen Secretary Treasurer Director Annika Farrell Warren D Allmon Trustees Samuel T Pees (to 6/30/95) Richard E Petit (to 6/30/96) Gary Rosenberg (to 6/30/96) James E Sorauf (to 6/30/94) John Steinmetz (to 6/30/94) Tucker Abbott (to 6/30/96) Bruce M Bell (to 6/30/96) Carlton E Brett (to 6/30/95) R Ann F Budd William (to 6/30/94) Crepet (to 6/30/94) J Thomas Dutro, Jr (to 6/30/96) Annika Farrell (to 6/30/95) Robert M Linsley (to 6/30/95) Peter McLaughlin (to 6/30/95) L Susan B Stephens (to 6/30/96) Henry W Theisen (to 6/30/95) Raymond Van Houtte (to 6/30/94) BULLETINS OF AMERICAN PALEONTOLOGY and PALAEONTOGRAPHICA AMERICANA Warren D Allmon Editor Reviewers for this issue Arthur Boucot J J Thomas Dutro, Jr A list of titles in both series, and available numbers and volumes may be had on request Volumes 1-23 of Bulletins of American Paleontolog}' have been reprinted by Kraus Reprint Corporation, Route 100, Millwood, New York 10546 USA Volume of Palaeontographica Americana has been reprinted by Johnson Fifth Ave., New York, NY 10003 USA Reprint Corporation, 1 1 Subscriptions to Bulletins of American Paleontology may be started at any volume or year Current price is US $50.00 per volume Numbers of time, by Palaeontographica Americana are priced individually, and are invoiced separately on request for additional information, write or call: Paleontological Research Institution 1259 Trumansburg Road Ithaca, NY 14850 USA (607) 273-6623 FAX This paper meets the requirements (607) 273-6620 of ANSI/NISO Z39.48-1992 (Permanence of Paper) Begun VOLUME 107, in 1895 NUMBER 346 JUNE Brachiopods of the Onondaga Formation Moorehouse Member (Devonian, in the Genesee Valley, Western Eifelian), New York by Howard R Feldman Paleontological Research Institution 1259 Trumansburg Road New York, 14850 U.S.A Ithaca, 15, 1994 ISSN 0007-5779 ISBN 0-87710-432-8 Library of Congress Catalog Card Number: 94-65488 Printed in the United States of America Allen Press, Inc Uwrence, KS 66044 U.S.A CONTENTS Page Abstract Introduction Acknowledgements Stratigraphic setting Systematic paleontology Introduction Philosophical considerations A Note on the use of open nomenclature Terminology Abbreviations of repository institutions and Measurement abbreviations and subscripts Systematics Appendix: Localities cited References cited Plates Index localities 9 in this report 36 36 42 50 LIST OF ILLUSTRATIONS Text-figure Index p^g^ map of collecting localities in the Onondaga Limestone Generalized stratigraphic column for the Onondaga Limestone LIST in the Genessee Valley OF TABLES '^^^^^ 10 1 12 14 16 17 19 20 22 23 24 Page Measurements of Dalejina cf aha (Hall, 863) Measurements of Schizophoria cf muhistnata Hall 859-1 86 Measurements oi Pentamerella arata (Conrad 1841) Measurements of Leptaena sp Measurements of " Brachyprion" cf mirabilis (Johnson, 970) Measurements of Brachyprion'! sp Measurements of Slrophodonla dernissa (Conrad, 842) Measurements of Coslislrophonella punctulifcra (Conrad, 1838) Measurements of Longispina mucronala (Hall, 1843) Measurements of "Eodevonaria" cf heinispherica (Hall 1857) Measurements of Cupularostrum? sp Measurements of Atr^pa "reticularis" (Linnaeus, 767) Measurements of Coelospira Camilla Hall 1867 Measurements oi Athyns boucoti n sp Measurements of Athyris leoni, n sp Measurements of Aferislina cf nasula (Conrad 1842) Measurements of Pentagonia unisulcata (Conrad 1841) Measurements of Nucleospira venlncosa (Hall 1857) Measurements of Tremalospira gibbosa Hall, 1859 and T camura Measurements of Alaliformia? sp Measurements of Megakozlowskiella raricosta (Conrad, 842) Measurements of Elylha fimbnala (Conrad, 842) Measurements of Cyrtma hamillonensis (Hall, 1857) Measurements of Cryptonellal sp ( 10 ) 12 13 15 15 17 17 19 19 21 1 (Hall, 1850) 22 23 24 25 26 29 29 30 31 33 34 35 35 BRACHIOPODS OF THE ONONDAGA FORMATION MOOREHOUSE MEMBER (DEVONIAN, EIFELIAN), IN THE GENESEE VALLEY, WESTERN NEW YORK Howard R Feldman* Biology Department New Touro College New York 10010 York, ABSTRACT In the Genesee Valley of western brachiopods, described herein m New Of the 26 York, the Moorehouse Member of the Onondaga Limestone contams 46 species of species of brachiopods in the underlying Bois Blanc Formation in western New York, and the Onondaga Formation; that is, taxa found in the Bois Blanc-Onondaga mterval mdicatc a high degree of stasis During Onondaga time there was a progressive increase in relative water depth throughout the basin as indicated by a gradual northward shift of the carbonate facies as well as a northward migration of the overlying Hamilton Group The Nedrow Member represents a minor transgressive cycle due to an influx of mud from the east, and the Moorehouse Member to Seneca Member interval represents a major transgression Brachiopods in the Moorehouse Member include Charionoides and Penlagonia genera endemic to the Appohimchi Subprovince Alhbonium halli and Discomyorthis'! sp are the only species herein not previously reported from Onondaga strata in western New York Two new species are erected, Athyris boucoli and A leoni The most significant change in brachiopod faunas across the state during Moorehouse time is the increasing abundance of stropheodontids towards the west 1 also occur the Onondaga and show no evolutionary change between INTRODUCTION in the As a continuation of my previous studies of Onondaga brachiopod taxonomy and paleoecology (Feldman, 1980, 1985; Feldman and Lindemann, 1986; Lindemann and Feldman, 1981, 1987; Racheboeufand Feldman, 1990) I have sampled and described the brachiopod fauna in the Genesee Valley of western New York The outcrop belt of the Onondaga New York extends from Port Jervis (Tris- (Text-figure Limestone in ) tate area) in the southeastern part of the state, north- eastward to Kingston and the Helderbergs, and then westward towards Syracuse, Rochester and Buffalo All of the brachiopods studied were recovered from the Moorehouse Member (see section on stratigraphic set- dense limestone with little shale Two methods of collecting were employed: Firstly, weathered fossils were collected from extensive, but rather rare, bedding surfaces in abandoned quarries and, secondly, blocks of limestone with silicified specimens were recovered, processed, and etched in an acid bath Approximately 180 kg of limestone yielded 190 silicified shells The brachiopod fauna of the Onondaga Limestone is particularly challenging to study because collecting is almost impossible unless bedding planes are accessible Vertical roadcut faces are often not useful as faunal identification is difficult, and outcrops with well-silicified shells are not always exposed As ting), a hard, * Also, Department of Invertebrates, The American Museum of Natural History 79th Street at Central Park West 10024 (correspondence to this address) New York N.Y it mid-Hudson is sporadic and from poor to very Valley, silicification the degree of silicification ranges good This study improves our understanding of brachiopod abundance and evolution in the Lower Middle Devonian, provides data for ecological and biogeographical studies, and improves correlations with more westerly limestone suites such as the Detroit River Group (Anderdon Limestone, Lucas Dolomite, Amherstburg Dolomite, Sylvania Sandstone) of the Michigan Basin, southwestern Ontario and north-central Ohio Previous paleocommunity and stratigraphic studies of the non-reefal aspects of the Onondaga Limestone (Oliver, 1954, 1956) have resulted in a fairly good understanding of the formation in southeastern New York (except for the area between Ellenville and Port Jervis where outcrops and complete sections are sparse) and central New York (Syracuse) The area around Rochester (i.e Genesee Valley), however, has not been studied in detail until now ACKNOWLEDGEMENTS I thank Arthur J Boucot (Oregon State University, Corvallis, Oregon) for his continued assistance, en- couragement, and support over the past decade, as my studies of the Onondaga Limestone have progressed from the mid-Hudson Valley, westward across New York State He reviewed the manuscript and made numerous valuable suggestions for improvement Touro College contributed $500.00 toward publication of this paper Bulletin 346 Nedrow, Moorehouse and Seneca A brief description of the lithologies of these members is provided here, but detailed stratigraphic descriptions may be found in Oliver (1954) —\n western New York the Edgea massive, light-gray, coarse, crystalline lime- Edgecliff Member cliff is stone about m thick, packed with solitary rugose and tabulate corals which form biostromes in many places Typical of the Edgecliff are large crinoid columnals and stems up to about 2.5 cm in diameter Near Buffalo the Edgecliff undergoes a facies change with at least one large, lens-shaped, biohermal ("reef) structure, which contains extremely irregular bedding (Oliver, 1954, p 635) Oliver (1954, p 636) also reports the occurrence of small "micro-reefs" exposed in the bioText-figure I — Index map of collecting localities in the Onondaga Limestone [Moorehouse Member] Genesee Valley, western New York A, Loc 52; A large exposure on the floor of eastern part of an abandoned quarry Lower Moorehouse Member; B, AMNH AMNH Loc 54; Numerous exposures at comer of an active Company Lower Moore- the southwest quarry operated by the Penfield Dolomite AMNH house Member; C, Loc 3153; Small exposures along the southeast part of an active quarry operated by the General Crushed Stone Company; Upper Moorehouse Member (for exact locations see Appendix) The University of Rochester (Rochester, New York), Gordon C Baird, State University College at Fredonia (New York) and George C McCarlton E Brett, Museum and Science Center (RochYork) spent valuable time with me in the and helped immensely in interpreting Onondaga intosh, Rochester ester field, New stratigraphy in western New York State Paul Copper, Laurentian University and J Thomas Jr., U.S Geological Survey (Washington, D.C.), Dutro, deserve thanks for script pod him critical comments on the manu- New York State Museum and Sci(Albany, New York), made the brachio- Ed Landing, ence Service collections in Albany available for study; I thank Fred Collier, (U.S National Museum of Natural History, Washington, D.C.) enabled me to examine the collections under his care and kindly loaned me specimens from the National collection in Washington Andrew Modell and Susan Klofak (both of the American Museum of Natural History, New York), deserve thanks for photographic work and specimen preparation, respectively Laura Lynne Gallo, Sarah Lawrence College (Bronxville, New York) graciously assisted me in the field and during my examination of the collection in Albany, New York for his hospitality STRATIGRAPHIC SETTING Five strome at Clarence, 20 km east of Williamsville In both western and eastern New York, bioherms occur 2), Onondaga, representing favorable conditions for and crinoids The Edgecliff is the growth of corals thought to be a moderately high energy, shallow-water, shelf carbonate which, in the western part of the state, becomes a crinoidal grainstone ( Woodrow et al 989) Clarence Member —The Clarence 11.5m thick, interfingers with the Edgecliff in places and essentially underlies the Nedrow Member in western New York Some workers believe that the Clarence replaces the Nedrow in the western part of the state (Ozol, 1963) The extremely cherty Clarence (up to 75% by volume; April et al 1984) consists of both vertically and hor- izontally coalescing chert nodules enclosing fine-grained lime mudstones (Selleck, 985) and is easily recognized in outcrop The rate of clastic deposition in the Clar- ence exceeded that of the Edgecliff, as indicated by the volume of clays The Clarence Member rep- greater resents slightly deeper water deposition than the Edge- most of the silica was derived biogenically from dissolution and reprecipitation of sponge spicules and, possibly radiolaria (Selleck, 1985) Accoringto Woodcliff; row et al belt enriched (1989) the Clarence in siliceous may represent a facies organisms m Nedrow Member —The Nedrow, 13 thick, is a dark-gray highly argillaceous limestone which forms recessed ledges when extensively weathered Fresh cuts, as at Jamesville, New York (AMNH Loc light- to 28) not have the row In eastern argillaceous crop, even New and when is typical appearance of the NedYork, the Nedrow becomes less more difficult to recognize in out- extensively weathered The Nedrow thought to represent deeper and muddier water deposition than the rest of the lower part of the formation and probably represents a minor transgressive cycle is in associated with an influx of elastics from the east (Woodrow f/ al 1989) the Edgecliff, Clarence, Moorehouse Member —Typically uniformly bed- members of the Onondaga Limestone occur the study area (Text-figure in the New York Devonian Brachiopods: Feldman m thick, is a medium-gray, Hmestone with abundant darkDuring Moorehouse time, there ap- ded, the Moorehouse, 1 fine-grained micritic weathering chert parently were shallowing conditions similar to those that prevailed during Edgecliff-Clarence time (Kissling and Moshier, 1981) The top of the Moorehouse is marked by the occurrence of the Onondaga Indian Nation metabentonite (= Tioga * TIOGA BENTONITE B metabentonite) which UNIFORMLY BEDDED WITH CHERT THROUGHOUT is not always present in the various quarries visited Most of the brachiopods studied in the Onondaga in the last decade were recovered from the Member Seneca Afeftiher.— The base of (4 m thick in western New is a medium- fossils the Seneca Member marked by the cm thick) The Seneca York) "Tioga B" ash layer (about 15 Moorehouse of the due, in part, to accessibility is to dark-gray, light-weathering wacke- stone, sparsely fossiliferous, with occasional chert nodules throughout The "Pink Hallinetes Zone" (= Zone layer and 1.8 m with chopacked thick, is a thinly bedded limestone pink At which are stained netid brachiopods many of most localities in western New York, a bed of chert can be found about 15 cm below the top of Zone J The upper part of the Seneca is more argillaceous and distinctly darker in appearance and is capped by a bone J of Oliver, 1954), m above the ash SHALE THROUGHOUT PARTICULARLY NOTICEABLE WHEN WEATHERED bed making the contact with the overlying basal Hamilton Group (Oatka Creek Shale) sharply defined in most places During Onondaga time there was a general increase in water depth throughout the basin as indicated by a gradual northward shift of all carbonate facies comprising the successive members and by northward migration with time of the Marcellus Shale (Kissling Moshier, 1981) According to Woodrow et al -•- I SHARP BREAK ^ ^^\ I j ^ ^ 1 VERY HACKLY WHITISH CHERT APPEARS ALMOST BRECCIATED and (1989), Moorehouse-to-Seneca stratigraphic succession a major transgression, with the upper Moorehouse and lowermost Seneca aerobic facies passing upwards into a dysaerobic, deeper water upper Seneca facies which is finally succeeded by the mini- the represents BUFF COLORED LIMESTONE WITH SCATTERED CORALS; NON-CHERTY MICRITIC mally dysaerobic to anaerobic Union Springs environ- -r ment KEY SYSTEMATIC PALEONTOLOGY Philosophical Considerations goals of this project is to help refine our understanding of the "invasion" of European taxa into North America by providing data for a biogeographic study of the Early Middle Devonian faunas in New York Species recognition is an integral part of eastern this work 10 DARK CHERT Introduction One of the SCALE A biological definition of a species is: a group of populations which replace each other geographically or ecologically and of which the neighboring ones in- T-T-n SS5 TTT xS — LIGHT CHERT LIMESTONE SHALE Text-figure daga Limestone — Generalized stratigraphic m the Genesee Valley column lor the Onon- Bulletin 346 tergrade or hybridize wherever they are in contact or which are potentially capable of doing so (with one or more of the populations) in those cases where contact is prevented by geographical or ecological barriers (Mayr, 1976) In other words, the modem biological concept of a species is based on an organism's having the potential to interbreed and produce fertile offspring in natural conditions (Guensburg 1984) The biological definition is untestable using fossils and, species have therefore been reported in this study based on morphology (i.e morphospecies) in the sense of Hoover (1981) Although the concept of a species may and sometimes does differ among paleontologists depending on the group studied and the preservation of the particular collections on hand (Sohn, 983), I consider, as Cooper and Dutro 982), species names merely conveniences for discussing clusters of morphologic variability within what are, for the most part, well1 ( Taxonomic names are, Hoover "handles" for discussion as (1981) has noted, of a definite group of specimens, and their correspondence to genetic groups becomes increasingly vague as established generic concepts the temporal distance from the Recent increases I con- "lumper" nor a "splitter." Of the two new species erected herein, one is based on 45 specimens while the other is based on four specimens As Raup and Stanley (1978) have so aptly stated, the fact that species discrimination depends largely on the experience of the person making the discrimination has led to an informal definition of the species that is invoked with surprising frequency: "A species is a species if a competent specialist says it is." After studying the morphology of the new taxa I have decided that they indeed represent new forms and not belong to any previously named species sider myself neither a A Open nomenclature is a device whereby an author own ma- expresses his or her judgement of his or her (Matthews, 1973) It is the procedure by which a taxonomist comments upon the identity of a specimen that cannot be readily or securely determined (Bengston, 1988) The use of "cf " before a speciesgroup name indicates a provisional determination for the species Here, "cf " stands for confer, not conforniis, and means "compare to" and not "compare with" (sensu Bengston, 988) I follow Bengston's ( 988) conterial "compare to" expresses a possible identity, which is what most taxonomists have in mind when they use "cf ", whereas "compare with" cept that the wording rather implies a distinction Further, "cf " is inserted between the genus and species name as in Levenea cf subcarinata and not Levenea cf L suhcarinata as advocated by Lucas (1986) As noted by Bengston (1988), the former expression conveys in an unambiguous way the message that the author considers the specimen in question to be "probably or possibly the species subcarinata although there is not enough material to be sure, but if it is subcarinata it should be referred to the genus Levenea.'' The use of "aff." indicates a new, undescribed taxon and relates it to a named taxon (sensu Bengston, 988) The use of"? sp." indicates an uncertain identification and, that specific identification specimens at is not possible with the hand because of poorly preserved present or original material Terminology The morphologic terms used herein follow the glosbrachiopod volume of the Treatise on In- sary in the vertebrate Paleontology (Williams et 1965, pp H139-H155) Note on the Use of Open Nomenclature Abbreviations of Repository Institutions The Code of Zoological is to promote stability names of animals, and intent of the International Nomenclature (StoU et 96 ) and universality in the scientific to insure that each name is unique and distinct Ac- and Repository of AMNH: freedom of taxonomic thought or action Matthews (1973) noted that although the Code sets a limit on its provisions, it does not in any way intend to impinge on the individual taxonomist's exercise of his or her judgement Since the Code provides no explicit guidelines for the use of "open nomenclature," I make the following brief comments regarding my use of it in this monograph Additional insight into the question of open nomenclature may be found in Bengston (1988), Komicker (1979), Matthews (1973), and Richter (1943, 1948) York, to these ends, all its and none restricts the [unnumbered] specimens unless otherwise indicated: provisions are subservient cording to the Code, all Localities American Museum of Natural History, New New York AMNH Loc: American Museum of Natural History number MCZ: Museum of Comparative Zoology, Harvard University, Cambridge Massachusetts NYSM: New York State Museum and Science Service, locality Albany, New York University of Michigan Museum of Paleontology, Ann Arbor, Michigan LTSNM: United States National Museum of Natural History, Smithsonian Institution, Washington, D.C UMMP: Bulletin 346 46 Explanation of Plate Figure 1-5 sp Pedicle, brachial, lateral, posterior, anterior Camarospira? 6-15 Alhyris boucoti new AMNH 14-15 Pedicle, brachial interior, new 44229 AMNH AMNH 44227 44228 (paratype), x4, AMNH Loc 3153 x2.5 AMNH Loc 3153 (paratype), AMNH Loc 3152 AMNH 44230, xl AMNH 44231, x from AMNH Loc 3152) AMNH 44232, x AMNH 44233, x 3, AMNH Loc 3153 interior, 1 (all 26 26 nasula (Conrad 1842) 25 Pedicle intenor, 26 Pedicle interior P^ge 23 25 24 Athyris sp A Brachial interior AMNH AMNH 27-31 Pedicle extenor, interior, 32-34 Menstina! Loc 3153 (holotype) x 2, 18-21 Pedicle exterior, posterior, pedicle, brachial interior 22-23 Brachial exterior, cf AMNH species 16-17 Pedicle interior, exterior, 25-31 Menstina 44226, x2 24 10-13 Pedicle, brachial, antenor posterior views 6-23 Athyris leoni AMNH species 6-9 Pedicle, brachial, anterior, posterior views views sp Pedicle extenor, 44234, x2 44235, x2 from AMNH AMNH 44236, x 1.5 AMNH Loc 52 AMNH 44237, x lateral, anterior, posterior postenor anterior, (all 2, Loc 3153) 27 Bulletins of American Paleontology, Volume 107 Plate Bulletins of American Paleontology, Volume Plate 107 ' ''iWw J •I '%2» W'^^i 10 12 13 L^ 14 New York Devonian Brachiopods: Feldman 47 Explanation of Plate Figure Mehstinal sp Pedicle interior, exterior, 3-6 Chanonoides dons (Hall 1860) 1-2 3^ Pedicle, brachial exterior, 5-6 Pedicle, brachial exterior, AMNH AMNH AMNH 44238, x 1.5, AMNH Page 27 27 Loc 3152 44239, x2.5 44240, x3 (both from AMNH Loc 3152) 7-13 Pentagonia unisulcata (Conrad, 1841) 28 7-10 Pedicle brachial, anterior, posterior views, 11 Pedicle intenor, AMNH AMNH 44241, x2 44242, x2 12-13 Brachial exterior, interior, AMNH 44243, x3 (all from AMNH Loc 3152) 14-23 Nucleospira venlncosa (Hall, 1857) 28 AMNH 44244, posterior, anterior views, AMNH 44245, AMNH 1.8, AMNH 14—17 Pedicle, brachial, posterior, anterior views, x2.5, Loc 3152 18-21 Pedicle, brachial, x Loc 3153 .AMNH 22 Pedicle interior, 23 Brachial interior, 44246, x4 44247, x5 (both from AMNH AMNH 24-33 Tremalospira gibbosa Hall, 1859 24-28 Pedicle, brachial, lateral, anterior, posterior views, 29-33 Pedicle, brachial, anterior, posterior, lateral views, Loc 3152) 29 AMNH 44248, AMNH 44249, x3.5 x3 (both from AMNH Loc 3152) Bulletin 346 48 Explanation of Plate Page Figure 1-4 Acrospirifer 7-9 30 (Hall, 1843) AMNH 44250, x 3^ Brachial exterior, intenor, AMNH 44251 x2 (both from AMNH Loc 3154) AMNH Loc 3154 MucrospiriferJ sp Brachial exterior, intenor, AMNH 44252 x posterior (x2) AMNH 44253 AMNH Alatiformial sp Pedicle exterior x 3), anlenor x 1-2 5-6 duodcnana Pedicle exterior, inlenor, 31 1, ( 10-15 Mediospirifcr sp 10-1 ( 2), Loc 3154 A 31 31 AMNH 24454 x 1.5 AMNH 44255 x 1.5 AMNH 44256 x 1.5 Brachial exterior, interior, 12-13 Pedicle exterior, interior, 14-15 Pedicle exterior, interior, (all 16-20 Mediospirifer? sp B Pedicle, brachial, from lateral, anterior, posterior AMNH views, Loc 3154) AMNH 44257, x AMNH Loc 3154 32 32 21-26 Megakoztowskietta raricosia (Conrad 842) 44258 xl 21 Posterior AMNH 22 Pedicle intenor AMNH 44259 x 23 Pedicle intenor AMNH 44260 x 24 Pedicle intenor, AMNH 44261, x2 25 Brachial intenor AMNH 44262, x 1.5 26 Brachial intenor, AMNH 44263, x I.5 Paraspinfer'^ sp Brachial exterior AMNH 44262 (all 27 AMNH Loc 3154) AMNH Loc 3153 from x2 33 33 28-32 Etytha fimbnala (Conrad 1842) AMNH 44265, AMNH Loc 3153 28-31 Pedicle, brachial, posterior, anterior views 44266, x2 32 Brachial intenor, AMNH x 1.5 AMNH Loc 3152 Bulletins of American Paleontology Volume 107 Plate Bulletins of American Paleontology, Volume 107 Plate * j*^ 1''.' ' t^t iằ>-,>.* 'W **- -i^- '>iô^ijib^ 13 >% / 16 '"1* 12 ' %'Ji^ J id^ 14 '^v, 15 New York Devonian Brachiopods: Feldman Explanation of Plate Page Figure 1-3 Etytha fimbhata (Conrad, 1842) 33 AMNH 44267, x3, AMNH Loc 3153 Brachial interior, AMNH 44268, x6.5, AMNH 3154 Detail of ornament (note spines) on pedicle exterior, AMNH Ambocoeliid indet Pedicle, brachial, anterior views, AMNH 44270, 4-6 49 Pedicle interior, AMNH 24469, x2, Loc 3152 x5, Loc 3152 AMNH 7-10 Cyrtina hamihonensis (Hall, 1857) AMNH 24471, x2.5 AMNH 24472, x3.5 (both from AMNH Loc 10 Brachial interior, AMNH 24473, x5, AMNH Loc 3153 Cyrtma sp A Pedicle exterior, AMNH 24474, x2, AMNH Loc 3152 34 34 7-8 Pedicle, brachial extenor, 11 Pedicle exterior, 3152) 35 35 12-17 Cryplonellaf sp AMNH 44275, x2.3 AMNH 44276, x3.7 (both from AMNH AMNH 44277, x3, AMNH Loc 3152 12-14 Pedicle, posterior, brachial views, 15-17 Brachial, postenor, pedicle views 18-19 CranaenaP sp Brachial, pedicle views, Loc 3152) 36 Bulletin 346 50 INDEX Note; Page numbers are in light lfTOip/r//er face; plate numbers Helmbrecht and Wedekind, 1923 murchisoni aff atlamicus (C\aTV.e 1907) ^Korfffiana Boucot and Johnson, 1968 duodenana (HM 1843) »7!!t;/(; Schuchcn and Cooper, 1932 Lindcmann and Feldman (1981) Lindcmann and Feldman (1987) "Chonetes" lincata, Chonetes lingtia Penlainerelta Idelined] Modell, liiicala (afV.) A 18 Moose River SyncUnonum 19 miicronala 13 18 3,18-19 15 Lochkovian 11 Longispina Cooper 1942 emmeiensis (WtnchcW) 1866 18 mucionalus Delthyris 18-19 M ucrospinfer Graban Lucas (1986) 3,75-79,44 Strophomena Carolina (HaW 1867) formosa mamensis Boucot, 1973 20 20 20 20 3,20,44 sp Schizophona 1-12 10 30 murchisoni Acrospirifer 31 Orthis 16 macrocosta Cupiilaroslntm 21 Museum magnapleura Megakozlowskwlla 33 20-21,29,31 1,1 Trematospira musculosa Orthis mamensis Machaeraha 5,31 Schizophoria 20 Mame 31 1931 multistriata Bowen 1967 whiltingloni 18 31 Mucrospirifer? sp (cf.) Machaeraria Cooper 1955 14,21,29,31 Chonelcs Longispina Llandoverian niiicronala (HaW 1843) 53 Michigan Ann Arbor Michigan Basin f-'agerholmi iohnson, 1970 A'av/o//a Johnson, 1970 sp Brachiopods: Feldman 10 of Comparative Zoology, Harvard University nasula Atrypa maria Merislella 20 26 Maryland 26 5,26-27 cf Mcristina 14 Mcnslella 26 Massachusetls, Cambridge Mcristina 26-27 Matthews (1973) Mayr(1976) McMonnigal Limestone navicula, Levenea Nevada 11 nevadensis Schizophoria mcdialis Delthyris 31 New New Mediospinfer^ sp B \fediospinfer Bubiichenko 1956 atidaculus (Conrad 1842) foniaculus (HaW 1857) sp A 7,i2,48 31 32 32 7,i7-i2,48 Megakozlowskiella Boucox 1957 cf rancasto Johnson, 1970 32 New Jersey, Scotland &O B 33 32 Bronx ville rancosta (Conrad 1842) 7.i2-ii,48 Caledonia HaW 1867) Megaslrophia? sp Megaslropliia Caster 1939 33 raricosla ( Megastrophiella Mcnslella ) mana HaW 1863 donsHaW 1860 IcntiformisCXarke 1900 /la^MM (Conrad, 1842) pnnceps(HaW 1857) j//;;i!(/ruw (Conrad) Nettlcroth 1889 Menstclla"^ umsulcata (Conrad) HaW 1862 Merisiinal sp Mcnslina HaW 1867 cf nasula (Conrad 1842) haskmsi (HaW 1860) toa Hall, 1859 nasula Boucot and Johnson, 1968 30 30 6,8 12,26,34 36 Railroad tracks 5-6 33 27 Buffalo Canandaigua Lake Central 19 16 Cherry Valley 34 16 Clarence 26 27 27 26 27 Danen 17 Delphi 25 2,76,43 Mcnslella (Pcntagonia) unisulcala (Conrad) Hall, 1867 11 26 Brunswick Formation Limestone New York Albany Albany [County] 32 magnaplcura io\\r\sor\ 1970 ranaMto Boucot and Johnson, 1968 15,18,30-31,33 28 28 28 5,6,27,46-47 26 5,26-27,46 27 27 26-27 Ellcnville Fredonia Fultonham Genesee [County] Genesee Valley Glenene Green Pond Outlier Gulf Road Hamilton Hclderberg Mountains Honeoye Honeoye Hudson 7,36 5-7,9-14,21,23,28-29,31,34-36 27,30 11,14,23,31 36 25 33 36 Falls Falls 33 Road 36 30 Bulletin 346 54 34 peersi Cozzens, 1846 36 unisulcata (Comad 1841) 30 unisiilcala (Conrad) Sa\a$e 1930 5,10.14,16.21,23-25,27-29,31.33-35 i/H/s»/cato (Conrad) Stauffer 1915 Knox Le Roy Lockport mid-Hudson Valley Monroe [County] Perry Road 36 36 Port Jervis 5-6 Rochester Rondout 27.30 34 Saugerties Southeastern Staflbrd Syracuse New York State Nikiforova et al (1985) nobilis Callipleura 13 pericosia Penlamerella 13 perofrata Tremalospira 30 32 6.8 lingiia Imbrie, perlamellosus Spirifer "Pink" Chonetes Zone "Pink" Hallinctcs Zone PUcostropheodonia SoV.o\%kaya, 1960 25 Plicoslropheodontal sp 21 Podolia Port Nucleospira atf venlricosa Boucot, 1973 aff ventricosa Fe\dman 1983 Boucot and Johnson 1968 venlricosa {HaW 1857) venlncosa HaW 1859 sp nuculoidea Alhyris 29 28-29 28-29 6.28-29.41 28 25 5.34 Oklahoma praecursor Protathyris Pragian Spirifer princeps Merislella Prololeploslrophia? sp Onondaga Indian Nation metabentonite Onondaga Limestone Members (cf 6-7 Lower Upper 6 Nedrow Member Seneca 5-7,36 Member "Onondaga" Limestone Ontario Oregon 30 27 14 2.14.43 punclulifera Coslislrophonella ) Coslilsrophonella punclulifera 3,17.75 17.18 17 Sirophodonia 17 Sirophomena 17 p.v [defined] Racheboeufand Feldman (1990) 5.19 raricosla (cf ) Megakozlowskiclla 5-6 Dellhvris 5-7 Megakozlowskiella 31 25 14 (cf.), 6-7 27,30 25 Member Member Moorehouse Member 2,/6,43 Prolalhyris praecursor Ko7-\owsV.i, 1929 Oliver (1956) Edgecliff 16 prohfica Schuchcrlella 20 5-7 Clarence 19 11 pnmacYus Oliver (1954) 5-7,20,24-25 13 25 Ewen Limestone Prololeploslrophia CasXer 1939 Ohio 12 33 6.28.33 Science Service 12 tumida \mbrie 1959 36 Museum and araM Boucot and Johnson, 1968 cf 13 l,2./2.42-43 11.14.23 5-7.12.14.20,22-23.26-27.31.33,35 Williamsville arara Hall, 1867 28 12-13 13 U.S Route 15 Western araw (Conrad, 1841) 28 1959 pericosia Imhne 1959 12.14.26.30.33 Schoharie [County] Penlamerella HaW 1867 ajionensis Imbrue 1959 28 6,28-29.47 (.\fegakozlonskiella) Kozlowskiella 5,25 Spirifer Raup and Stanley (1978) 32 31 7,32-33 32 32 Corvallis 5.25 Recent State University 5.25 reclicoslalum Cupularoslrum 21 reclirostra Terebratula 35 Oriskany Sandstone 30,34 Orlhis alsus HaW 1863 murchisoni D'Archiac and de VemeuW 1842 Muscn/osa HaW 1857 Siibcannala HaW 1857 Ozol (1963) 16 reimanni Cryplonella 35 Anomia "relicularis." Alrypa 22 relicularis 4.22 10 Rhipidomella a/.va Hall 1867 rhomhoidalis Slenoscisma 21 10 "rhomboidalis" parafragilis Schizophoria 11 paraprhna, Schizophoria 11 7.ii,48 Paraspirifer? sp Paraspinfcr VJedekind, 1926 aciiniinauis (Conrad 1839) peersi 33 28 Pentagonia Penfield Dolomite 33 Company 6.36 Leplaena Leplaena "Rhynchonella" Carolina Hall 1867 Rhynchonella fonnosa HaW 1857 Richter(1943) Richter(1948) Rochester Museum and Science Center (afT.) 13 (cf.) 13 Pennsylvania 14 Rochester Shale Pentagonia Cozzens 1846 28 romingeri lenla(Ha\\, 1867) 28 Cranaena 20 20 8 30 36 New York Devonian Brachiopods: Feldman Terchralula 36 "Schucherlella" nigosa Leptacna 13 Schucherlella'? Rutgers University 26 55 14 14 Spinocyrlla "curyleines" (Ov/en, 1844) 32 Splrlfer alaliformi.s Sallisaw Formation 20 Schizophoria King, 1850 10 cf multislrmta cf multislnata Feldman 1985 ci muhistriala (HaW 1859-1861) cf Paraprima 1-12 \ 10-11.42 ferronensis \mbrie 1959 12 DTQvcTnrann 1907 /;m/)//u/u Hall spiriferoides Alhyris II St parafragilis iohnson 1970 II State University College at Fredonia 12 S'?c«a5rw/»a /w/// Fagerstrom 1961 traversensis Grabau, 1931 33-34 Schoharie Grit Schucherlella pro/Z/zca Schuchert, 1913 "Schucherlella" hecraftensis (Clarke 900) sp.B Johnson 1970 sp Feldman, 1985 sp A Boucot and Johnson 1968 A Boucot Gauri and Southard 1970 B Boucot and Johnson 1968 schucherli Laurent Limestone (Hall //!()/Ji/)
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