(CTC^ -American OLUME 102, JUNE NUMBER 340 Neogene Paleontology 14 in the northern 26, 1992 Dominican Republic Otoliths of teleostean fishes by Dirk Nolf and Gary L Stringer Paleontological Research Institution 1259 Trumansburg Road New York, 14850 U.S.A Ithaca, ^ PALEONTOLOGICAL RESEARCH INSTITUTION Officers Harry A Lefrngwell J Thomas Dutro, Jr Henry W Theisen James C Showacre Roger J Howley Peter R Hoover Henry W Theisen President Vice-President Secretary Treasurer Assistant Treasurer Director Legal Counsel Trustees Edward Bruce M Bell (to 6/30/93) Carlton E Brett (to 6/30/92) William L Crepet (to 6/30/94) Thomas Dutro, Jr (to 6/30/93) Harry A Leffingwell (to 6/30/93) J Robert M Linsley (to 6/30/92) Samuel T Pees (to 6/30/92) William P S B Picou, Jr (to 6/30/92) Constance A Sancetta (to 6/30/94) James C Showacre (to 6/30/93) James E Sorauf (to 6/30/94) John Steinmetz (to 6/30/94) Henry W Theisen (to 6/30/92) Raymond Van Houtte Ventress (to (to 6/30/94) 6/30/93) BULLETINS OF AMERICAN 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the following individuals and institutions PATRONS ($1000 or more at the discretion of the contributor) Allen (1967) American Oil Company (1976) Atlantic Richreld Company (1978) Christina L Balk (1970, 1982 1983) Hans M Bolli (1984) James E Ruth G Browne (1986) Mr & Mrs Kenneth E Caster (1967) Chevron Oil Company (1978 1982) Exxon Company (1977 to date) Lois S Fogelsanger (1966) Gulf Oil Corporation 1978) Merrill W Haas (1975) ( Robert C Hoerle (1974-1977) Richard I Johnson (1967, 1986) J M McDonald Foundation (1972 1978) Mobil Oil Corporation (1977 to date) Samuel T Pees (1981) Richard E Petit (1983) Robert A Pohowsky (1982) Texaco, Inc (1987 to date) Union Oil of California (1982 to date) United States Steel Foundation (1976) Charles G Ventress (1983 to date) 976-1 984) Christine C Wakeley ( (continued overleaf LIFE MEMBERS ($400) R Ralph Tucker Abbott Langenheim, L James E Allen Elizabeth A Balcells-Baldwin Christina L Balk Bruce M Bell Harry Robert A Black Richard S Boardman Donald Egbert G Leigh, Gerard Jr Lenhard A Louie N Marincovich, Moore R Shuji Niko Hiroshi Noda Hans Bolli David John Bottjer Ruth G Browne J David Bukry Sy3IL B Burger Sakae O'Hara William A Oliver, Samuel T Pees Richard E Ptiii Lyle D Campbell Edward Carter Anneliese S Caster Kenneth E Caster John E DuPont J Thomas Dutro, Jr Robert A Pohowsky John Pojeta, Jr John K Pope Anthony Reso Arthur W Rocker John J L Mark Erickson Richard J Erickson Lois S Fogelsanger A Eugene Fritsche Christopher L Garvie Ernest H Gilmour Merrill W Haas Anita G Harris Steven M Herrick Carole S Hickman Robert C Hoerle F D Holland, Jr Frederick H C Hotchkiss David Jablonski Richard \ Johnson David B Jones Peter Jung Tomoki Kase Jr A Leffinowell B Picou, Jr Arnold Ross Walter E Sage, John B Jr III Saunders Judith Schiebout Edward S Slagle Robert E Sloan Richard L Squires David H Stansbery Jorge P Valdes Raymond Van Houtte Charles G Ventress Wiluam p S Ventress Emily H Vokes Harold E Vokes Christine C Wakeley Thomas Waller R Patricia H Kelley Albert D Warren, Jr Gary D Webster Ralph H Willoughby David Garrett Kerr Armour Cecil H Kindle WiLLL\M F Klose Thomas JiRi Kriz II Winslow Yancey Zullo C E Victor A Jr OLUME 102, NUMBER 340 JUNE Neogene Paleontology 14 in the northern Dominican Republic Otoliths of teleostean fishes by Dirk Nolf and Gary L Stringer Paleontological Research Institution 1259 Trumansburg Road New York, 14850 U.S.A Ithaca, 26, 1992 Library of Congress Card Number: 85-63715 Printed in the United States of America Allen Press, Inc Lawrence, KS 66044 U.S.A CONTENTS Page 45 45 46 48 48 48 Abstract Resumen Introduction Acknowledgments Abbreviations of Repository Institutions Some anatomical and morphological remarks on otoliths Systematic Paleontology 49 Introduction and remarks on taxa requiring comments Genus Ptotosus Lacepede, 803 Genus Diaphus Eigenmann and Eigenmann Description of new species Genus Genus Genus Genus 51 51 Brotula Cuvier, 1829 51 Lepophidium Gill, 1895 Otophidium Gill in Jordan, 1885 Neobythiles Goode and Bean, 885 52 52 53 Tribe Dinematichthyini "genus Dinematichthyinorum" Genus Apogon Lacepede, 80 Genus Lactarius Valenciennes in Cuvier and Valenciennes, 1833 Genus Conodon Cuvier in Cuvier and Valenciennes, 1830 Genus Haemulon Cuvier, 1829 Genus Clenosciaena Fowler and Bean, 1923 Genus Larimus Cuvier in Cuvier and Valenciennes, 830 Genus Mentiarrhus Gill, 1861b Genus Ophioscion Gill, 1863 Genus Pareques Gill m Goode, 1876 1 Family Gobiidae Bonaparte, 1832 Family Pleuronectidae Rafinesque, 1815 Conclusions Critical considerations of the available data 53 54 54 54 55 55 56 56 56 56 56 57 57 Paleoenvironmental evaluation 57 and relationships of the Paleogene and Recent Caribbean fauna Comparison with other Neogene otolith associations in the Eastern Nonh American and Caribbean realm Appendix Alphabetical list of otolith-based fish taxa in the localities from the Neogene of the Dominican Republic 60 References Cited 63 66 76 Paleobiogeographic Plates Index affinities 61 61 LIST OF ILLUSTRATIONS ''^^e Text-figure of the Cibao Valley, northern Dominican Republic Position of otoliths in the braincase ot Oncorhynchus mykiss (Walbaum, 792), and in the labyrinth oi Salvelinus fontinalis (Mitchill, 46 1814) 'S Geological sketch map Morphology of the inner face of a saccular otolith ''9 LIST OF TABLES Page Table of the river valleys studied for this project, showing approximate correlations List of teleost species represented by otoliths in the Neogene of the Dominican Republic Present-day bathymetric range of taxa represented in the Neogene of the Dominican Republic Stratigraphic sections from six 47 50 58 NEOCENE PALEONTOLOGY IN THE NORTHERN DOMINICAN REPUBLIC 14 Otoliths of teleostean fishes by Dirk Nolf' Gary and Stringer^ L ABSTRACT The study of area, northern fossil otoliths has allowed us to reconstruct a teleost fauna of 84 species for the Neogene of the Cibao Valley attributed to nominal species and 53 are described in open Dominican Republic Thirty-one of those could be Recent Central American species or very closely related species, and Lepophidium latesulcaliim Otophidium robinsi Olophidium robustum, "genus Dinematichthyinorum" stmlhmmzi, "'genus Dmematichthyinorum" sauli Conodon moreauxi Clenosciaena latecaudata, and Menliarrhtis chaoi Many taxa could not be identified at the species level because the knowledge of otoliths of -.elated Recent Central American species is too restricted Although the bulk of the material comes from the late Miocene NNl Zone, restricted associations from other levels allow us to draw some paleoenvironmental conclusions for most of the levels The association from the Baitoa Formation consists of very shallow-water taxa that all occur in depths of less than 50 meters The associations from the Cercado Formation and from the the basal part of the Gurabo Formation consist essentially of neritic taxa that lived of the taxa are also regular inhabitants of euryhaline environments such as lagoons in depths of less than 50 meters At least or estuaries Beside these forms, some deeper-ranging taxa also occur, but none of these are confined to soft-bottom neritic environments The information available for the NN12-NN13 interval of the Gurabo Formation is very scattered The otolith associations are restricted in the Mao Formation (NN14-NN15 interval), but it is possible to conclude that the environment nomenclature Among the nominal species, 22 belong to eight represent extinct species that are new; 1 was deep neritic or, more likely, upper slope The studied fauna can be considered very close to that of the present-day Caribbean, except for Plotosus L^acepede, 1803 and Lactarim Valenciennes in Cuvier and Valenciennes, 1833, which we interpret as relicts of the Paleogene Western Tethys fauna In terms of both abundance in number of specimens and taxonomic diversity, the bestrepresented groups are ophidiids, haemulids, and sciaenids These groups are also well-represented in Paleogene otolith associations of the U S Gulf Coast and in the Recent Caribbean fauna Analysis of the available data provides evidence that near the Oligocene-Miocene boundary, important evolutionary events affected some of the groups of Caribbean teleosts, and that in the concerned groups, affinities of the Neogene forms are much closer to the Recent fauna than to the Paleogene one A weak point in such a statement however, is our complete ignorance of Paleogene faunas for the southern Caribbean and eastern Pacific realm, because the possibility exists that in these areas relatives of "modem" Central American taxa already existed in the Paleogene RESUMEN de otolitos fosiles nos permitido reconstruir una fauna teleostea de 84 especies en el Neogeno del Valle Cibao, el none de la Repiiblica Dominicana De estos taxones, treinta y uno han sido atribuidos a especies nominales y 53 descriptos nomenclatura abierta Dentro de las especies nominales, vientidos pertenecen a especies centroamericanas recientes, o especies consanguineas, mientras que ocho representan nuevas especies extintas; Lepophidium latesulcaliim, Otophidium robinsi Otophidium robustum, "genus Dinematichthyinorum" smithvanizi, "genus Dinematichthyinorum" sauli, Conodon moreauxi, Clenosciaena latecaudata, y Menticirrhus chaoi Muchos taxones no han podido identificados a nivel especifico porque el conocimiento de otolitos de especies centroamericanas consanguineas recientes es demasiado restringido Aiin cuando (Mioceno tardio), asociaciones restringidas de otros niveles nos fwrmiten la mayor parte de las muestras provienen de la zona N llegar a conclusiones paleoambientales para la mayor parte de los niveles Las asociaciones de la Formacion Baitoa consisten en taxones de aguas muy someras localizadas a menos de 50 m de profundidad Las asociaciones de la Formacion Cercado y de la parte basal de la Formacion Gurabo consisten esencialmente de taxones neriticos que vivieron en profundidades menores que El estudio localizado en 50 m Por lo menos de los taxones son tambien habitantes regulares de ambientes salobres como albiiferas o estuarios Ademas de estas formas, tambien se encuentran algunos taxones que habitan aguas profundas, pero ninguno de estos se halla confinado a ambientes neriticos de sedimentacion fina La informacion disponible de las zonas NN 2-NN de la Formacion Gurabo se encuentra muy dispersada Las asociaciones otoliticas estan restringidas a la Formacion Mao (zonas NNI4-NN15), pero es posible concluir que el paleoambiente fue neritico de aguas profundas, o, probablemente, de la parte alta del talud continental Se puede considerar a la fauna estudiada muy cercana a la actual del Mar Caribe, excepcion de Plotosus Lacepede, 1803 y Lactarius Valenciennes en Cuvier y Valenciennes, 1833, que interpretamos como relictos de la fauna paleogena del Tethys occidental En terminos de abundancia de especimenes y de diversidad taxonomica, los grupos mejores representados son ophidiidos, haemulidos, y sciaenidos Tambien estos grupos se hallan bien representados en las asociaciones otoliticas paleogenas 1 Royal Belgian Institute of Natural Sciences, 29, Vautierstraat, 1040 Brussels, 110 Patton Drive, West Monroe, LA 29 U S A 1 , BELGIUM Bulletin 340 46 de la Mexico de costa del golfo de los Estados Unidos, como asi tambien en la fauna reciente del Mar Caribe Analisis de los datos disponibles muestran evidencias de que cerca del limite Oligoceno-Mioceno, importantes eventos evolutivos afectaron a unos de los gnipos de teleosteos del Caribe, y dentro de estos grupos, las formas Neogenas se parecen mas a las faunas modemas que a las del Paleogeno Un punto debil en esta declaracion es, sin embargo, nuestra falta de conocimiento acerca de las faunas paleogenas de las regiones del sur de Caribe y este del Oceano Pacifico porque existe la posibilidad de que en estas areas, parientes de estos taxones centroamericanos "modemos" hubieran existido en el Paleogeno INTRODUCTION Otoliths, or ear stones, are very common fossils of and are present in a broad spectrum of sedimentary environments They are unique among teleostean fishes, vertebrate fossils in the sense that they are not parts of the skeletons but specialized hard parts of the acoustico-lateralis system The mineral composition of otoliths is calcium carbonate, not calcium phosphate like in bone In most genera, otoliths are species-diagnostic and represent a valuable tool in the reconstruction of the teleostean fauna of a specific geologic period In contrast to the situation in Europe, little is known North American Tertiary The first report on North American otoliths is by Koken (1888), who described 23 species from Paleogene deposits of the U S Gulf Coast Except for some casual remarks (Eastman, 1904), no otolith-based faunal reconstruction was published before the work of Frizzell and Dante (1965), who described 24 species resulting from an eclectic sampling of the U S Gulf Coast Paleogene Previously, Frizzell and Lamber (1961, 962) published two short articles, one on "myripristids" and about the otoliths Text-figure in the — Geological sketch map of the Cibao Valley, northern (Saunders, Jung and Biju-Duval, 1986, text-fig 3, p 7) one on congrid otoliths from the same area and age Several species described in the above cited papers appeared in faunal lists of two localities in the late Eocene of Louisiana (Stringer, 1979, 1986), but no formal descriptions of additional species were included The above cited publications, except for Eastman (1904), deal with Paleogene material and although they are not directly related to the present study, provide information on the Paleogene history of teleosts in the Caribbean and Gulf Coast areas For the Neogene of the U S Gulf and Atlantic coasts, only two relevant papers treat fossil fish otoliths: Fitch and Lavenberg (1983) provided a list of 45 otolithbased taxa, including Recent species, from the Pliocene of the Lee Creek Mine, North Carolina; Clarke and Fitch (1979), in a paper on cephalopods, referred to the presence of otoliths of ten fish families in the early Miocene Chipola Formation, about 80 km east of Tallahassee, Florida In the Plio-Pleistocene of California, the presence of many Recent eastern Pacific species has been reported in various publications by Fitch and collaborators, but those data not pertain to the present topic Dominican Republic, showing areas from which samples were collected Dominican Republic Neogene 14: Nolf and Stringer Explanation of Plate 71 Page Figure Centroprislis philadelphica {Lmnaeui, 1758) IRSNB, nght otolith, inner view, off Georgia coast, U S comparative purposes] A., Recent [included for 62 Pristigenys species NMB P87 bis, right otolith (a, inner view; b, ventral view), loc TU 1227A, Arroyo Zaiaya, turbidity flow lens Pliocene, NN14 Zone 3-5 Centropristis species 3, NMB 4, NMB Pliocene lens Diplectrum species Epinephelus species TU 1227A, Arroyo Zaiaya turbidity flow NMB 1591 1, Rio Gurabo, Cercado Formation, Gulf of Mexico offTexas coast, late NNl Miocene 2^ne Recent [included for comparative purposes] loc TU 1230, Rio Cana, Cercado Formation, late Miocene otolith, inner view, off Haiti, Recent left 62 inner view, loc TU inner view, loc NMB 16854, Rio Cana, Cercado Formation, otolith, inner view, loc NMB 15907, Rio Gurabo, Cercado Formation, NMB 17268, Rio left otolith, Apogon species NMB P93, 1227A, Arroyo Zaiaya, turbidity flow lens Pliocene NN14 Zone 54 left otolith, late Miocene Epigonus species NMB 13 left P95, 62 late Miocene NNl Zone 54 left otolith, Apogon species NMB P94, Apogon species NMB inner view, P92, loc 62 left otolith, Epinephelus striatus {Bloch 1792) NMB P89, right otolith, inner view, otolith (a, inner view; b, ventral view) left IRSNB, loc P91, NMB Zone P90, right otolith, inner view, Diplecirum bhittatum (Valenciennes, 1828) NMB NN14 5, IRSNB, 61 P88, nght otolith; inner view; inner view, loc Yaque del Norte, Gurabo Formation, Pliocene NN12 Zone 54 P96, right otolith, inner view, loc NMB 17003, Rio Cana, Cercado Formation, late Miocene Lactarius species NMB 54 P97, right otolith (a, inner view; b, ventral view), loc NMB 15904, Rio Gurabo, Cercado Formation, late Miocene NNl Zone 15, 16 Apogon 15, late 54 species NMB P98, left otolith, inner view; 16, NMB P99, right otolith, inner view, loc NMB Miocene 17 Luljanus campechanus Poey, 1861 18 IRSNB, left otolith, inner view Gulf of Mexico, offTexas, U Ocyunts chrysurus (B\och 1791) IRSNB, 19, 20 20, left otolith (a, inner view; b, ventral view), off Haiti, S A., Recent [included for comparative purposes] Recent [included for comparative purposes] Ocyurus aff O chrysurus (Bloch 1791) 19 16837, Rio Cana, Cercado Formation, NMB NMB Lutjanus NMB Zone PlOO, left PlOl, left otolith, aff P102, otolith (a, 62 inner view; b, ventral view), loc inner view, loc TU 1294, Rio NMB 16837, Rio Cana, Cercado Formation, late Miocene Mao, Cercado Formation, late Miocene NNl I Zone campechanus Poey, 1861 L left otolith (a, inner ^iew; b, ventral view), loc 62 NMB 16923, Rio Mao, Cercado Formation, late Miocene NNll Bulletin 340 72 Explanation of Plate Page Figure Pristipomoides aquilonaris (Goode and Bean, IRSNB, left otolith, 896) inner view, off Mobile, Alabama, U S A., Recent [included for comparative purposes] 62 2-4 Pristipomoides species NMB PI 03, inner view; left otolith, Zalaya, turbidity flow lens Pliocene 4, NMB P105, right otolith NMB NN14 PI 04, left 12, 13 NMB NMB NMB Gurabo Formation, 62 late Miocene NN 62 PI 07, right otolith, inner view, P108, left otolith, inner view, 9, Eucinostomus species PI 09 right otolith (a, NMB 17288, Rio Yaque del Norte, Baitoa Formation, early or middle Miocene NMB 16935, Rio Yaque del Norte, Baitoa Formation, early or middle Miocene loc loc 62 inner view; b, ventral view), loc TU 1359, Rio Gurabo, Cercado Formation, late Miocene NNl Zone Anisotremus davidsoniH,Sxein6^chnsT 1875) IRSNB left otolith (a, inner view; Moharra rhombea (Cuvier, 1829) 13, NMB NMB NN 1, Pomadasys species 12, 16910, Rio Mao, Maury's Bluff NMB 11 NMB loc IRSNB, left otolith, inner view, near Fortaleza Brazil, Recent [included for comparative purposes] fuc/noiromuj Aavana (Nichols, 1912) IRSNB, left otolith, inner view, Bermuda, Recent [included for comparative purposes] Eucinostomus species 15910, Rio Gurabo, Cercado Formation, PI 06, nght otolith (a, inner view; b, ventral view), loc Zone 8, 1227A, Arroyo Zone inner view; b, ventral view), NMB 8, TU otolith (a, inner view; b, ventral view), loc Miocene NNl Zone Eucinostomus lefroyi {Goode 1874) late (a, 3, left PI nght otolith 1, ventral view), off California, U S A., Recent [included for comparative purposes] otolith, inner view, loc PI 10, b, (a, 62 TU 1294, Rio Miocene NNl Zone 15904, Rio Gurabo, Cercado Formation, Mao, Cercado Formation, NMB inner view; b, ventral view), loc late late Miocene Zone 61 Anisotremus species NMB PI 12 left otolith (a, inner view; b, ventral view), loc NMB Haemulon steindachneri QotAan and G\\ben 1883) IRSNB, left otolith, inner view, off Guyana, Recent [included 16-19 Haemulon species 17005, Rio Cana, Cercado Formation, late Miocene 15 16, 18, NMB NMB NMB PI 13, left otolith, inner view; 17, NMB PI for comparative purposes], 55 14, left otolith, inner view; 19, NMB 15903, Rio Gurabo, Cercado Formation, late Miocene NNl Zone 16923, Rio Mao, Cercado Formation, left otolith, inner view, loc PI 15, right otolith, inner view; P146, NMB late Miocene NNl Zone loc Bulletins of American Paleontology, Volume 102 Plate 14 -^ '2 ^3a Pristipomoides aquilonaris (GOODE8iBEANJ896) Eucinostomus lefroyi Pristipomoides sp (GOODE,1874) Eucinostomus havana (NICHOLSJ912) Motiarra rhombea {CUVIER,1829) I 1mm I 10a Eucinostomus sp.1 Eucinostomus sp.2 Anisotremus sp 1mm Anisotremus davidsonii Pomadasys sp (STEINDACHNER,1875) Haemulon sp l-laemulon steindachnerl (JORDAN 8,GILBERTJ882) Bulletins of American Paleontology, Volume 7a Plate 15 02 \^^— _._._.,«-^-'i>-'8 Lonchopislhus micrognathus (POEY,1861) 1mm 1mm , 1mm Mugil sp Polydactylus sp 1mm Opistognathus lonchurus JORDAN &GILBERT,1882 Mug/7 liza VALENCIENNES,1836 Polydactylus virginicus (LINNAEUS, 1758) R Dominican Republic Neocene 14; Nolf and Stringer Explanation of Plate 73 Page Figure , 4-6 Conodon nobilis (Linnaeus 1758) 1, IRSNB, left otolith (a inner view; b, ventral, view); 2, IRSNB, left otolith, inner view, Caribbean, Recent Conodon serrifer Jordan and Gilbert, 1883 IRSNB right otolith (a, inner view; b, ventral view) Gulf of California, Recent Conodon moreauxi, new species 4, NMB PI 16, holotype, NN Miocene 5, 6, 7-9 NMB NMB 1 left otolith (a, inner view; b, ventral view), loc NMB 55 55 54 15910, Rio Gurabo, Cercado Formation, late Zone otolith, inner view, loc PI 17, paratype, left PI 18, paratype, left otolith, inner view, loc NMB TU 16918, Rio Mao, Cercado Formation, late Miocene 1359, Rio Gurabo, Cercado Formation, late Miocene NNl NNl 1 Zone Zone Lonchopisthus micrognalhus (Poey, 1861) 7, NMB PI 19, 62 otolith (a, inner view; b, ventral view), loc left TU 1227A, Arroyo Zalaya, turbidity flow lens Pliocene NN14 Zone 10 1 8, IRSNB, 9, NMB IRSNB, left otolith Mugil species loc of Puerto Rico, Recent NMB PI 21, left (a, 15864, Rio Gurabo, Gurabo Formation, late Miocene NNl Zone inner view; b, ventral view), off Puerto Rico, Recent [included for comparative purposes] otolith, inner view, loc 62 TU 1278, Arroyo east of Rio Gurabo, Mu^// /iza Valenciennes, 1836 IRSNB, left otolith of juvenile specimen, inner vie v, H.iiti, Gurabo Formation, near Mio-Pliocene boundary Recent [included for comparative purposes] 62 Polydactylus species NMB 14 otolith, inner view, off coast 0/»/jWgna»Aui /oncAun/j Jordan and Gilbert, 1883 NMB 12 left PI 20, right otolith, inner view, PI 22, left otolith, inner view, loc TU 1282, Rio Cana, Cercado Formation, late Miocene Polydactylus virginicus (Linnaeus, 1758) IRSNB, right otolith, inner view, Margarita Island, Venezuela, Recent [included for comparative purposes] Bulletin 340 74 Explanation of Plate Page Figure and Valenciennes 1,2 Bairdiella ronchus (Cuvier in Cuvier IRSNB, NMB left P123, otolith, inner view, off left Ophioscion species Rio de Janeiro otolith, inner view loc TU 61 830) Brazil, Recent 1294 Rio Mao, Cercado Formation, late Miocene NNl Zone 56 NMB PI 24 nght otolith, inner view loc NMB 16913 Rio Mao, Maury's Bluff 3, Cercado Formation, late Miocene NNl Zone 5, NMB NMB 6, 7, P125, I NMB 17287, Rio Yaque del Norte, Baitoa Formation, early or middle Miocene 55 15903, Rio Gurabo, Cercado Formation, late Miocene NNl 15907, Rio Gurabo, Cercado Formation, late Miocene NN Zone 1 Zone 56 758) Margarita Island, Venezuela, Recent otolith, inner view, off 56 left otolith, left NMB inner view, loc 15904, Rio Gurabo, Cercado Formation, late Miocene NNl Zone 56 littoralis left otolith, PI 30, Caribbean Recent otolith, inner view (Holbrook, 56 860) inner view, off Rio de Janeiro, Brazil, Recent Larimus breviceps Cuvier left otolith, in Cuvier and Valenciennes, inner view loc NMB 56 830 15903 Rio Gurabo, Cercado Formation, late Miocene NNl Zone 56 Pareques species 14, left Menticirrhus 13, Zone Menticirrhus saxatilis (Bloch and Schneider, 1801) NMB 13, 14 NNl 56 otolith, inner view, loc PI 29, holotype, IRSNB, 15907, Rio Gurabo, Cercado Formation, late Miocene Menticirrhus chaoi new species IRSNB, left Menticirrhus americanus (Linnaeus, NMB NMB NMB PI 27 holotype left otolith, inner view, loc NMB NMB PI 28 paratype, left otolith, inner view, loc NMB IRSNB loc Ctenosciaena lalecaudata new species 6, P126, utricular otolith, inner view, Ophioscion species NMB NMB PI31 nght otolith, inner view, loc PI 32 right otolith, inner view loc Equetus lanceolatus (Linnaeus, BMNH, Stinton TU TU Gurabo Formation, Pliocene 1354, Rio Cana Canada de Zamba, Gurabo Formation, 1206, roadcut at K17, Pliocene 56 758) coll., left otolith, inner view, off Curasao, Recent 16 Pareques acuminatus (Bloch and Schneider 1801) LACM, Fitch coll left otolith, inner view, off Puerto Rico, Recent 56 17 Equetus punctatus (Bloch and Schneider 56 IRSNB, left otolith, inner view, Umbrina species 80 ) off Haiti, Recent NMB P133, left otolith, inner view, loc NMB 62 16932, Rio Mao, Maury's Bluff 2, Cercado Formation, late Miocene NNl I Zone Bulletins of American Paleontology, Volume 02 Plate 16 R,(F) 1mm Ophioscion sp.l 1mm Ophioscion sp.2 1mm Bairdiella ronchus (CUVIER,1830) 1mm Ophioscion sp Ctenosciaena latecaudata n.sp Menticirrhus americanus (LINNAEUS,1758) 1mm Menticirrhus chaoi n.sp 1mm Menticirrhus saxatilis (BLOCH & SCHNEIDER,1801) Equetus lanceolatus (LINNAEUS,1758) Menticirrhus littoralis (HOLBROOK,1855) 1mm 1mm Equetus punctatus Larimus breviceps CUVIER;830 Pareques acuminatus (BLOCH&SCHNEIDER,1801) (BLOCH&SCHNEIDER,180:) Umbrina sp Bulletins of American Paleontology, Volume 102 Plate 17 Dominican Republic Neocene 14: Nolf and Stringer Explanation of Plate 75 Page Figure Sphyraena species NMB 2, PI 34, 62 inner view, left otolith, loc NMB Gurabo Formation, 16818, Rio Cana, Pliocene Dactyloscopus byersi Dawson, 1969 LACM, Fitch coll 2, left otolith (a, inner view; b, ventral view); comparative purposes] Labrisomus species PI 35, left otolith (a inner view; b, ventral view), loc Dactyloscopidae indeterminate P136, right otolith (a, inner view; b, ventral view), loc 3, left otolith, inner view Gulf of California, Recent [included for 62 NMB NMB NMB TU 17005, Rio Cana, Cercado Formation, late Miocene 61 1206, roadcut at K17, Gurabo Formation, late Miocene 6-11 Nes longus (Nichols, 1914) IRSNB, 12 left otoliths, Gobiidae species P137, left otolith Zone Gobiidae species NMB NMB PI 38, 56 inner views, off Bermuda, Recent 57 inner view; (a, c, outer view), loc 16915, Rio Mao, Cercado Formation, late Miocene NNll 57 otolith (a, inner view; b, ventral view), loc left NMB NMB 15864, Rio Gurabo, Gurabo Formation, late Miocene NMB 15823, Rio Gurabo, NNl Zone 14 Gobiidae species P139, left otolith Bothidae indeterminate NMB 15, 16 15, 16, 7, NMB PI 40, left 57 (a, inner view; b, ventral view), Zone NMB PI41, right Mao Formation, Pliocene NN15 Zone 61 otolith (a, inner view; NNll loc c, otolith, inner view, loc posterior view), loc NMB NMB 16924, Rio Mao, Cercado Formation, late Miocene 16923, Rio Mao, Cercado Formation, late Miocene NNl I Zone Cynoglossidae indeterminate 61 NMB PI 42, left otolith (a, inner view; c, posterior view), loc NMB 59 2, Rio Gurabo, Cercado Formation, late Miocene NNl I Zone 18, 19 62 Soleidae indeterminate 18, 19, NMB NMB PI 43, left otolith, TU 1294, Rio Mao, Cercado Formation, late Miocene NNl Zone Zone loc TU 1373, Rio Gurabo, Cercado Formation, late Miocene NNl inner view, loc PI 44, right otolith, inner view, 1 57 20 ? Pleuronectidae indeterminate NMB PI 45, right otolith, inner view, loc TU I227A, Arroyo Zalaya, turbidite, Pliocene NNl Zone Bulletin 340 76 INDEX Note: Page numbers are in light face; plate numbers are in bold face type; the page 49 actinopterygian saccular otolith, morphology of acununatiis Pareques Ne-umia aequalis, aff 16 11 50,59,62,69 Myclophum affine Scopoh (e.v 49 68 10 Gronow), 1777 59,61 sp 50.6/.67 50 Albulidae American Museum of Natural History, New York, New York, americanus, Menticirrhus 16 angnillahs Plotosus 10 \ SI 5) 14 14 sp ANSP [Academy of Natural U.S.A 63 56,74 57,68 A nisol remits davidsonii (Steindachner, 59 72 50,67,72 Sciences, Philadelphia, Pennsylvania, U.S.A.] 48,57 Antilles Sea 70 Apogon Lacepede, 1801 54,59,61 48,74 Bohlke and Robins (1959) Bohlke, Eugenia indet brachycephalus (af[.), breviceps 13 50,54,61,71 Brotula Cuvier 1829 49,50,61 Ariidae Ariosoma ba/earica {Delaroche IS09) 50.59,67,67 selcnops Reid \934 a.([ .-Ir/Mj 9 A selenops Reid \9i4 59.67 50,58,67,67 Valenciennes /« Cuvier and Valenciennes, 1840 58,59 10 sp Alherinomorus stipes (Miiller 50.67,68 and Trolschel, 1847) 50,59,67,70 Atlantic Ocean 56 54 Queensland Australian realm 51 49.56,60 Bairdiella G\\\, 1861a ronchus (Cuvier m Cuvier and Valenciennes 1830) 50,59,67,74 16 Baitoa Formation balearica Ariosoma 45,50,52,55-58,72,74 50,59,67,67 barbata Brotula 12 /ia/-/)a/a 70 59 brasiliensis] 12 50.62,70 51,53,55 50,68 58,59 50,67,69 50 Larimus 16 50,56,59,62,74 57,59,60 (Bloch and Schneider, 1801) 51,52 clarkae Hnbbs 1944 51 Hubhs 1944 11 w«/n/)ar/)a/a (Temminck and Schlegel 1846) ordwayi Hildebrand and Barton, 949 50,57,61.69 sp 50,57,61,69 aff B clarkae 51 52 11 lownsendi Vov/\eT 1900 52 Brolulina Fow\eT 1946 sp 54 54 of Nolf(1980) 48 Brzobhaty, Rostislav 17 Dactyloscopus 75 50 Bythitidae 51,52 10 50,59,62,68 caribbaeus Coelorinchus 11 50,59,67,69 Caribbean region 59 Bahamas 50 48 Barbados Bavay, Daniel Hygophum Hygophum (aff.) 10 59,6S 50,62,68 10 60 Berg (1895) bermudensis Carapus bermudensis{aff.).Carapus Berra(1981) hivittatum Diplectrum Bleeker(1852) Bloch (1785-1795) Bloch and Schneider (1801) 59 50,67,69 11 13 campechanus Lutjanus campechanus (aff.) Luljanus Canary Islands Cape Verde Islands Carapidae Carapus bermudensis (Jones 1874) aff C bermudensis (Jones ISl 4) caribbaea Saurida Batrachoididae benoiti 50,57,62,68 11 bathyal taxa benoiii 57,59,68 46,52 western Australia 10 10 sp byersi, 12 Diaphus Herniramphus Hyporhamphiis [= Herniramphus brasiliensis (afT.), Herniramphus Brasso Formation Breder(1927) Bregmaceros Thompson, 1840 Bregmacerotidae Area beds 50,67,75 brasiliensis, 50,54,61,71 72 47 17 47,61 brachycephalns Diaphus 50.54.61.71 14 56 Bowden Formation 13 aquilonaris, Pristipomoides 58 50,57,61 13 50,54,61 48 Bothidae sp sp 53,69 Bold (1988) Bonaparte (1832) sp Apogonidae 74 Stinton Collection 50,54,61,71 italics BMNH [British Museum (Natural History), London, England, U.K.] 13 sp principal discussions occur are in 56,74 [defined] -l/te/a numbers on which Bermuda Cuba 13 59,77 50,62,71 68 69 50 59 11 50,67,69 45-47,49,51,52,54,55,57,60,61 70 54,61 57,72,75 Islands 67,68,70 Dominican Republic, Cibao Valley Arroyo Puiial Arroyo Zalaya [= Canada Zalaya] 45-47,57,58,60 46 51 13 71 58 50,51,68,71 51,56,74 46.47.50.53-56.58.67-73.75 Dominican Republic Neogene Guayubin 46 46 46 46 46 46 46 Janico Los Quemados Mao Moca Moncion Navarrete Rio Amina Potrero 47 RioCana 46,47,50,52,53,55,56,60,67-75 Canada de Zamba Rio Guayubin 74 46 46.47,50.52-56,60,67-75 RioGurabo Rio Mao Maury's Bluffl Maury's Bluff2 Maury's BluffS Rio Verde Rio Yaque del Norte Angostura Gorge Arroyo Hondo Arroyo Lopez 46,47,50,55,56,67,68,70-75 46,47.61 50 Clinidae Clupeidae 50,61 indet 50.68 Coelorinchus caribbaeus (Goode and Bean 1885) 55.71,73.74 47,54,61,69 73,74 45,60 47,51-56,60,61.70 Trinidad 45,48,49,54-57,60,61,73,74 12 easier (1958) easier (1966) 70 47 47,61 Cauda [defined] 49 Central America 45.47.51.52.55.56.60 70 61 Panama 61 50 59 Centropomidae Centropomiis 59 Centropristis nSS) 13 13 sp 71 50.6/, 71 45,47,50.52.53,55-59.67-75 Chao(1978) 45,50,55,61,74 16 latecaudala n sp per«v;a«a Chirichigno, 1969 Curasao Cuvier(1816) Cuvier(1829) Cuvier and Valenciennes (1828-1849) 55 74 60 50,51.55.60.72 45,50,54,56,58,60.61.74 50 Cynoglossidae 17 indet 50.67,75 60 Cvnoscion Gin, 1861b Dactyloscopidae 50,61 17 indet 50,67,75 Daclyloscopus byersi Dawson \969 17 davidsonii Anisotremus 14 Dawson (1969) 75 72 75 Delaroche(1809) Dermatopsis OpXby 1896 Diaphus Eigenmann and Eigenmann, 1890 brachycephalus T&ning 1928 aff D brachycepfialus Tinin% 192S 50.67 54 57,58.59 10 50.57,62.68 10 50,57.62.68 10 sp 50,57,62,68 5i, 54, 58, 60 Dinematichthyini "genus Dinematichthyinorum" sauli, n sp 12 "genus Dinematichthyinorum" 57,59,68 10 i»;;//n'a«;z/, n 12 45,50,5i,54,62,70 sp 45,49,50,54,62,70 59 Diplectrum 13 Wv/«afi/w (Valenciennes, 1828) sp 71 13 50,62,71 11 53,5i,69 49 dorsal run [defined] 56 N 56 chaoi Menlicirrhus 45,50,56,62,74 16 Chaunacidae Chaunaxsp chickarney 55 ^ra(:(7;arW!i« (Metzelaar 1919) 49 [defined] L 49 49 55.59,60 C/e^o.sc/ae^a Fowler and Bean, 1923 dormilalor Otophidium Cercado Formation 54.55.59 45,50,54,55,61,73 crista superior [defined] 50.6/ 70 12 sp 15 sp crista inferior [defined] sp Mexico Acapulco Veracruz n 55,73 Haiti Caribbean Sea carolinus Prionotus moreauxi, 55,73 Jamaica Puerto Rico southern 46,50,60,61 Conoiion Cuvier /« Cuvier and Valenciennes 1830 15 46 46 46 Zamba 48 48 49 15 46,47 Valverde 50,59,6/,69 11 M Cohen, Daniel Collete Bruce B coUicidum [defined] Congridae wi;fa (Linnaeus, 1758) 47 47 47 47 47 47 47 Santiago 50.67.67 Cocco(1838) iern/er Jordan and Gilbert 1883 46.47.50.53.67-72,74 Santiago Rodriguez 50,57,61,69 11 47.67,74 46,50.53,57.68-70 La Barranca La Boca Lopez Chao Brolula (aff.), Clarke and Fitch (1979) 47,68,72 47,74 Baitoa Cliff cf clarkae 77 46,47,50,52 Dam philadelphica (Linnaeus Nolf and Stringer 14: 50 50,6/,69 11 53 Otophidium 46.55,61 Chipola Formation 55 Chirichigno (1969) Ocyurus chrysunis (aff.), Ocyurus 13 chrysurus clarkae Brotula 13 59,77 50,62,71 51 46 Eastman (1904) Eigenmann and Eigenmann (1890) 51,58 50,61 Engraulidae mdet Epigonus sp 50,62,67 13 50,62,71 13 50,62,71 59 59 Epinephelus sp slriatus {Bloch 1792) epipelagic taxa 13 71 59 Bulletin 340 78 Equetus Rafinesque 1815 lanceolatus (Linnaeus \15S) pi//i'/ sp (Goode, 1874) 14 59 72 14 72 Giinther(1878) 50,69 Gurabo Formation 45,47,50,52-55.57.58.67-75 Haemulidae Haemulon 45 Cuvier, 1829 55,59,60 sp 14 sp sleindachneri (Jordan 14 50,62,72 12 50,62,70 Euleptorhamphus 59 sp iWavPoey, 1868 12 Europe France, southwest, Aquitaine Portugal euryhaline taxa excissura [defined] 14 50,62,72 70 54 54 54 59 49 and 50,55,62,72 Hartel Karsten Havana Eucinoslomus 14 Hemiramphidae 50 Hemiraniphus brasiliensis (Linnaeus \15S) 12 50,62,70 50 Hildebrand and Barton (1949) Hildehrandia flava (Goode and Bean, 1896) 52 Fitch flava 46,52,61 John 46,47 Hildehrandia fontinalis 50,59,62,67 70 48 Florida Strait Sahelinus Formalin® 55 52 Fowler (1900) Fowler (1946) Fowler and Bean (1923) Frizzell and Dante (1965) Frizzell and Umber (1961) Frizzell and Lamber (1962) 54 50,59,62,67 56,74 Holocentridae 50 HolocentrusScopoli(exGTonov/), 1777 58.59.61 12 sp 50.51.69 51 10 H benoili (Cocco \S38) hygomi Garman 50,62,68 68 10 (Liilken 1892) Hyporhamph us brasiliensis [= Hemiramphus ]M2) 59 hrasiliensis] aff // 12 Mmfef;a/w5(Ranzani 1842) 12 59,70 50.62.70 69 (1899) Gerreidae Gibbs, Robert 59,6S 10 unifascialus (Ranzani, 61 68 10 benoil i (Cocco ISiS) aff Gadoididae 50,62,70 Hubbs (1944) Hubbs and Schultz (1939) hygomi Hygophum Hygophum 55,60 46 46 46 70 12 H brasiliensis (Lmnaeus \75S) Heterenchelyidae aff Holbrook (1860) Fitch and Lavenberg (1983) 72 48 72 14 Gilbert, 1883) 50,61 H 48 indefatigable, Otophidium 53 indet Gill (1861a) 49,56,60 Bothidae 17 50.6/ 75 Gill (1861b) 56,60 Clupeidae 50.6/.67 Gill (1862a) 56,60 Cynoglossidae 17 50.6/ 75 Gill (1862b) 58 Dactyloscopidae 17 50 6/ 75 Gill (1863) 56,58 Engraulidae Gill (1895) 52,53 ? Pleuronectidae 47 Gillette (1984) gillii 12 Neohythites 50,5i,59,62,70 Girard (1854) 51 Globigerinoides trilobus fislulosus Globoroialia e.xilis Zone Zone Zone Zone Globorotalia rnargahiae Glororoialia miocenica 12 50.62.70 Soleidae 17 50.62,75 Indo-Pacific region Ireland 47,50 IRSNB 47,50 Gobiidae Bonaparte, 1832 69 Belgium] 50,57,62,75 50.57,62,75 Johnson, G David sp 17 50.57.62,75 Jones (1874) 72 Goode (1876) Goode and Bean Goode and Bean Goode and Bean gracilicirrhus, Greenwood 56,58,60 (1885) 50.53,58.69,70 (1886) 50.53,70 (1896) 50,58,67,72 Ctenosciaena et at (1966) Greenwood P Humphrey Gulf of California Gulf of Guinea Gulf of Mexico Gulf of Thailand 60 50,56,57,61 17 Goode(1874) 45,48,66-75 hopisthus GiW, 1862a 17 51,52,54 [Royal Belgian Institute of Natural Sciences, Brussels sp sp 50,62,67 50.57.62.75 Scorpaenidae 47,50 47.50.70 17 55 49 48 48 50,69 52 Jordan (1885) Jordan and Bollman (1890) Jordan and Gilbert (1883) 53 53,55,72,73 48 Jung, Peter kalHon Lepophidium 52 Koken (1888) 46 55,73,75 67 71 68 Labrisomus Sv/ainson 1839 sp Lacepede (1798-1803) 58.59 17 50.62.75 45,51,54.60.61 Dominican Republic Neocene LACM [Los Angeles County Museum of Natural History, 48,74,75 Los Angeles, California, U.S.A.] 74,75 Fitch Collection 50 Lactariidae LacfanMj Valenciennes /« Cuvier and Valenciennes, 1833 45,54,59,60 13 sp 50.5'#,62,71 lanceolatus, Equetus /.ar/ww5 Cuvier I/! Cuvier /)rei7fe/)i Cuvier and Valenciennes 1830 56,60,61 Cuvier and Valenciennes, 1830 //I lalecaudata Ctenosciaena latesulcatum 56,74 16 Lepophidium « a/ lefroyi kallion 51 46 52 48 Liem, Karl 50,55,56,60,70,71,73,74 70 56,74 73 15 1862b 58 micrognathus {Poey 1861) 15 lonchurus Opistognathus Nariva Formation 51 A'eMi Cuvier ;>i Cuvier and Valenciennes 1830 Neobythites Goode and Bean 1885 60 ^////VGoode and Bean 1885 ma»-g;>!a/!« sp Goode and Bean, 1886 12 50,5i,59,62,70 of Nolf (1976) 53 60 Neobythitini 59 neritic taxa /Vm/o«^m 17 (Nichols 1914) New Mexico 56,57,75 Bureau of Mines and Mineral Resources, Socorro, Nezumia New 48 Mexico, U.S.A aequalis {Gvmiher, \S1&) 50,59,62,69 11 Nichols (1912) 72 Nichols (1914) 56,75 56,57,75 17 50 nitidulum NMB Myctophum [Naturhistorishes 69 11 Museum Basel, Basel Switzerland] 48,52-57,61,62,66-75 Lutjanus campechanus Poty 13 \Z(>\ c-awpec/!a«i« Poey 86 59,7/ 13 50,62,71 68 Liitken (1892) Macrodon Sc\an7., 60 1822 Macrouridae 50,58 macrunis Pythonichthys Mao Formation Mao Adentro 67 Manzanilla Formation 55,56 45,47,50.53,57-59,68,69,70,75 Limestone nobilis Conodon 15 55,73 47,49,51-56 51-54 Nolf (1976) Nolf (1980) Nolf (1985) Nolf and Capetta (1989) Nolf and Lapierre (1979) Nolf and Steurbaut (1988) Nolf and Steurbaut (1989) Nolf, Dirk North America 49,50,53-56,60,61 51 55 51 51 54.60 47,61 47 marginatus Neobythites 50,5i, 59,62, 70 12 Ocyurus Mauritania 68 chrysurus {Bloch \79l} Mayer (1974) 58 aff MCZ 50,5i, 59 62,70 73 Lutjanidae aff / 5i,58 12 50,59,62,73 15 longus.Nes 48 Myripristidae 47,61 16 50,62,68 mykiss Onchorhynchus 58 52 Mugil 11 10 myriaster Porichthys 45,50,52,62,69 slaurophor Kob\n%, 1959a Gill, 59 68 69 10 72 Robms 1959b Lonchopisthus 51 50,51 48 60 Leriche (1938) liza multiharbata Brotula 45,50,52,62,69 11 50,70 Myctophidae Atyctophum a^«e(Lutken, 1892) 11 52,53,59 Linnaeus (1758) Linnaeus (1771) liltoralis Menticirrhus 50 and Trotschel (1845-1849) Mijller nitidulum Garman, \899 14 latesulcatum n sp 50,62,73 Mugilidae sp 1895 73 15 15 50,56.59,62.74 Eucinostomus GM Valenciennes, 1836 sp 45,50,55,61,74 Lepophidiini Lepophidium liza 68 16 (1980) 79 Morocco, Marrakech Mugil 16 La venberg, Robert Lee Nolf and Stringer 14: [Museum of Comparative Zoology, Harvard University, 48 Cambridge, Massachusetts, U.S.A.] Menticirrhus G'lW 1861b 56,59,60 amenca«wi (Linnaeus 1758) 56,74 16 chaoi.n.ip /;Hora/H(Holbrook, 1860) 16 45,50,56,62,74 ia\ar/7;5(Bloch and Schneider, 1801) 13 O Opistognathus lonchurus ]0Tdan and GiXbtn, ordwayi Brotula ostial rim [defined] ostium [defined] Orop/?;(/;»m Gill /n Jordan, 1885 c/;;fAan;f>r Bohlke and Robins, 1959 50,61 Xii^ii 50,62.67 73 15 52 49 49 52,53,59 53 Bulletin 340 80 Olophidium rfomiHawrBohlke and Robins, 1959 (/om;;Va/orB6hlke and Robins 1959 indefatigable ]oTdar\ and Bollman 1890 o/KOi/(^WM/M (Jordan and Gilbert, 1883) robimi.n.%p 11 robustum n 11 53,5i,69 11 53,5i,69 53 53 45,50,52,53,62,69 12 sp 45,50,5i,62,70 Pachypops GiU, 1861b Pacific Ocean, 60 sp Rafinesque ( Reid (1934) Rhechias lysanochila (Reid I9i4) Robins C Richard western 52 Pacific plate 51 Robins Ray and Douglass paracanthopterygian saccular otolith, morphology of 49 robinsi 56,58,60 acMmma?M5 (Bloch and Schneider, 1801) 16 sp Parophidion aff P schmidti 56,74 16 50,56,62,74 (Woods and Kanazawa 1951) 50,62,70 12 55 13 philadelphica Cenlropristis Gill, 71 60 1861b Pleuronectidae Rafinesque, 1815 50,57 17 ? Pleuronectidae indet 50,57.62,75 Plotosidae 45,57,54,59,60,62 a/!^«7/am(Bloch, 1794) 10 sp 5/,68 10 (1858-1861) (1866-1868) (1875-1877) 52 ( 1986) 45,50,52,53,62,69 11 12 45,50,5i,62.70 16 50,59,67,74 Rosen and Greenwood (1970) Rosenblatt and Rubinoff(1972) 48 58 49 rostrum [defined] Salvelmus fontinaiis (Unch\\\, 1814) Sample KR 1862 of Nolf (1976) 12 SauridacaribbaeaBKdir.\911 saxatihs Menlicirrhus 50,71.73 Schinz (1822) schmidti (af^.) Scopoli (1777) 58 Scorpaen idae Polyclemus Berg, 1895 60 59 selenops 50,62,73 selenops v;>g/«/cuj (Linnaeus 15 758) Polyipnus sp 50,62,72 Girard 1854 Hubbs and 57,59 Schultz, 1939 sp 51 50,57.62,69 11 posterior rim [defined] Prionotus caro/(«M.s (Linnaeus, 1771) 12 sp 70 12 50,62,70 13 50,62,71 Pristigenys 59 sp Prislipomoides Bleeker, 1852 (Goode and Bean, sp punclalus Equetus 58,59 896) 14 14 72 Pylhonichthys Poey, 1867 macrwnis (Regan, 1912) microphlhalmiis (Regan 1912) 59,67 50.58.67,67 50 15 55,73 60 58 63 48,54 smithvanizi "genus Dinematichthyinorum" 12 45,49,50,5'^,62,70 50 Soleidae 17 indet South America 50,62,75 47 56 Brazil 72 Fortaleza 74 Rio de Janeiro Colombia 61 French Guyana 69 72 68 Guyana Venezuela 70.73.74 Margarita Island sp., Anisotremus 14 56,74 Arius 10 50,67,68 58,59 Bregmaceros 11 50,67,69 50,62,72 16 Smith, C Lavett Smith-Vaniz, William F 49 50 Priacanthidae 50,62,70 Conodon 12 14 50 Ariosoma Ariosoma 60 sp 58,61 (aK.), Smith and Kanazawa (1977) 50 60 45,47,50,56,60,61 Serranidae serrifer 56,74 60 50,62,70 47,61 12 Sirembini 50,60,61 12 indet 59 50,62,70 Pomadasyidae Pomadasys \jactphde, 1803 50.59.62,68 16 Sciaenidae 50.62.68 Polymixiidae 46,47,50,57,58,61 10 Parophidion 10 Polymixia sp 73 59 45,50,5i,54,62,70 Schubert (1909) Sciaena Linnaeus, 1758 58 15 57 48,53 Saunders, Jung, and Biju-Duval( 1986) Poll (1959) sp 51 sample size [discussed] Saul William G sauli "genus Dinematichthyinorum" Poll (1954) Polydaclylus 48 Poll (1953) a^M/VonflTO 56 48,52,63 56-58 ronchus Bairdiella 58 58 «2iT;airer 50,62,72 52 Olophidium robustum Olophidium 50.5/,68 58 70 (1868) /'or/c/jr/iy^ 14 50 Plotosus LaceptAe 1803 Poey Poey Poey Poey 50,59,62,67 48 Patterson Colin peruviana Cienosciaena Plagioscwn 50,58,67 Moharra rhoinbea 60 Pareques Gin in Goode 1876 50,70 67 Regan (1912) southeastern 45,46,51,52,55 56,57 5) Ranzani (1842) Robins (1959a) Robins (1959b) Robins el al (1980) eastern 50,62,67 67 67 Alhula Brotula Brolutina 11 50.67.67 50,67,72 50,57,61,69 54 Dominican Republic Neocene 14: Nolf and Stringer Gary L 54,55.61 Centropomits 12 50.67,70 Stringer, Centroprislis 13 50,67.71 sulcus [defined] Chaunax 11 50,67,69 Diplcctruin 13 13 50,62,71 Swainson (1838-1839) Synodontidae 13 50.62,71 Tamana Formation 12 50,62,70 50,62,70 Tuning (1928) Temminck and Schlegel (1842-1850) Tethys 50,62,75 Thompson Epigonus Epinephetus Eiileplorhamphus Haemulon 50,62,71 50 55.62,72 14 Holocentnis Lahrisomus 50 54,62,71 13 Laclarius 50,62,73 Triglidae Myctophum 50,62,68 TU Neohythites 53 50,62,67 Opislhonema 51 45,51,54,60 58 (1840) 52 50,61 [Tulane University, New Orleans, Louisiana, U.S.A.] 47,48,52-59,61,62,67-75 tysanochila Rhechias 50,59,62,67 50 56,62,74 16 Plotosus 10 50,57,68 Polydaclylus 15 50,62,73 Polyipnus 10 50,62,68 Polymixia 12 50,62,70 14 Pomadasys 56 50,51,68 townsendi Brotula Miigil Pareques 49 58 50 t'»;/'n/!a 59.60 Cuvier, 1816 16 sp 50,62.74 UMML [University of Miami, Rosenstiel School of Marine and 48,53 Atmospheric Sciences, Miami, Florida, U.S.A.] Hyporhamphus Hyporhamphus 50,62,72 unifasciatus unifasctatus (aS.), 59,70 12 12 50,62,70 Prionotus 12 57,62,69 50,62,70 Pristigenys 13 50,62,71 Alabama Pristopomoides 14 50,62,72 Mobile Pylhonwhlhys Sphyraena 50,62,67 California 46,72 17 50,62,75 Florida 55,67 Umbrina 16 50,62,74 II Portchlhys sp 50, United States, Miami 13 50,54,61,71 Diaphus 10 50,57,62,68 Eucmoslomus 14 50,62,72 Gobiidae 17 50,57,62,75 Ophioscion 16 50,56,62,74 Apogon 13 50,54,61,71 Diaphus Eucinostomus 10 50,57,62,68 Gobiidae 17 50.57.62,75 Ophioscion 16 50,56.62.74 Apogon 13 50,54,61,71 Gobiidae 17 50,57,62,75 13 14 50.62,72 46 70,71 Georgia Gulf Coast 45,46,54,60,71 46 45 60 Louisiana West Monroe North Carolina Lee Creek Mine 46,61 Pennsylvania, Philadelphia 53,54 52,61 71 Texas 48 University of Brno, Czechoslovakia USNM sp 3, Apogon [United States National Museum of Natural History, Smithsonian Institution, Washington, DC, U.S.A.] 50,54,61,71 Valenciennes (1828) Sparidae 61 Valenciennes (1836) Sphyraena 59 velox 17 sp 50,62,75 Euleptorhamphus 50 Springvale Formation 51 virginicus, Polydaclylus 48 52 72 72 56 50 54 Vokes, Emily State University of staurophor, Ghent, Belgium Lepophidium Steindachner (1875) steindachneri Stellifer Haemulon OV.en (ex C\x\'\tv), 14 1817 Stemoptychidae Steurbaut (1984) Alhennomorus strialus Epinephelus Stringer (1979) Stringer (1986) 12 Walbaum (1792) Weiler (1959) 77 Weitzman, Stanley H Whitehead et al (1984-1986) Whitehead, Peter J P Woods and Kanazawa (1951) 46 46 Zidek, 50.59.67,70 13 73 12 70 49 49 15 73 48 48 ventral rim [defined] Vokes, Harold Jiri 48 71 ventral furrow [defined] Sphyraenidae stipes 61 Tallahassee Mississippi sp 2, '^ 52,63 northwestern 1, Apogon %X>- 60 72 48 47,61 48 58 48 50,70 48 ... numbers and volumes may be had on request Volumes 1-23 of Bulletins of ADierican Paleontology have been reprinted by Kraus Reprint Corporation, Route 100, Millwood, New York 10546 USA Volume oi... Crustacea) Bulletins of American Paleontology, vol 94, No 329, pp 1- 1958 figs., Delaroche, F figs W Copeia, pp 510-514, Naturelle, Paris, Annales, vol 13, 75 pp., pis Studies on fishes of the family... of the Fishes of the Eastern Coast of North America, from Greenland to Georgia Philadelphia Acad- Bulletin 340 64 emy of Natural Sciences, Proceedings for 1860, 63 pp Revision of the Genera of