... 20 Levin, S and Segel, L., Hypothesis for origin of planktonic patchiness, Nature, 259, 65 9, 19 76 â 2004 by CRC Press LLC 408 Handbook of Scaling Methods in Aquatic Ecology: Measurement, Analysis, ... Pattern-oriented modelling in population ecology, Sci Total Environ., 183, 151, 19 96 â 2004 by CRC Press LLC 4 26 Handbook of Scaling Methods in Aquatic Ecology: Measurement, Analysis, Simulation ... 445 â 2004 by CRC Press LLC 4 46 Handbook of Scaling Methods in Aquatic Ecology: Measurement, Analysis, Simulation not sinking particles, tended to collect in downwelling regions with a vertical...