... Clément and Janin [5], the predominant occurrence of mineral inclusions in heartwood offers an objective standardto discriminate sapwood from heartwood rings, even in specieswith indistinct or ... pixels and minimum, maximum and average surface proportion (in %) of suchpixels in the X-rays of 0.7 mm and 2 mm strips of trees I, II and IV(60 rings analysed per strip).0.7 mm 2 mmTree Min ... formation of mineral inclusions isinduced or increases when the main functions of the woodytissues shift from transport and storage of water, minerals and assimilates, as in sapwood and earlywood,...